scieee Science in your language
[en] (orig)

Extrapolating base-line trunk shrinkage reference equations across olive orchards

Abstract

Maximum daily trunk shrinkage is a common measurement in irrigation scheduling of fruit trees. But the strong relationship between these measurements and the environment severely limit field applications. Reference baselines are the solution for understanding the influence of environmental conditions. Nevertheless, the extrapolation out of the original conditions is not clear. The aim of this study was to compare several approaches to estimate a reference baseline in an olive orchard where there were no previous data from other seasons. Two orchards, separated 60. m, with different tree density were used. Orchard 1 had greater tree density than orchard 2, though the age and the cultivar were the same. Trunk diameters of both orchards were similar but the crown volume of orchard 2 was slightly lower than orchard 1. The current reference baselines of maximum daily trunk diameter in both orchards were not significantly different between them (p<0.05). In orchard 1, the previous reference baseline was calculated in a 5-year study (the so called multi-seasons approach). The multi-seasons approach was not significantly different in slope but it was in the y-interception to the current reference baselines in both orchards (p<0.05). This approach over-estimated the values in both orchards. Two additional approaches were tested. These latter approaches used data before massive pit hardening to estimate the current reference baseline. One of them used the early data to estimate a complete reference baseline (the so-called early approach). The other (the so-called y-early approach) used the same data only to estimate the y-interception and assumed that the slope was the same as in the multi-seasons approach. The early approach under-estimated the value of maximum daily trunk shrinkage. The early-y approach provided a satisfactory estimation of the reference baseline and improved those obtained with the multi-seasons approach. The limitations and uses in irrigation scheduling are also discussed.

Read accessible full text

Extrapolating base-line trunk shrinkage reference equations across olive orchards

Author: Corell González, Mireia; Girón Moreno, Ignacio Francisco; Moriana Elvira, Alfonso; Dell'Amico, José; Morales, D.; Moreno Lucas, Félix
Publisher: Elsevier
Year: 2016
DOI: 10.1016/j.agwat.2013.04.011
Source: https://idus.us.es/bitstreams/41c01707-5f56-46fe-8b3c-2f2d53577692/download
1"
"
Ex apola ing base-line unk sh inkage e e ence equa ions ac oss oli e o cha ds 1"
. 2"
3"
Co ell, M.a,*, Gi ón, I.F.b, Mo iana, A.a, Dell’Amico, J.c, Mo ales, D.c, Mo eno, F.b 4"
5"
aEscuela Técnica Supe io de Ingenie ía Ag onómica. Uni e si y o Se ille, Ca e e a 6"
de U e a Km 1, 41013 Se illa, Spain 7"
bIns i u o de Recu sos Na u ales y Ag obiología de Se illa (IRNAS-CSIC), P.O. Box 8"
1052, E-41080 Se illa, Spain. 9"
c Ins i u o Nacional de Ciencias Ag ícolas, Cuba 10"
11"
*Co esponding au ho : [email p o ec ed] Phone: (+34)954487298; Fax: (+34)954486436 12"
13"
14"
15"
16"
17"
18"
19"
20"
2"
"
Abs ac 21"
Maximum daily unk sh inkage is a common measu emen in i ia ion scheduling o 22"
ui ees. Bu he s ong ela ionship be ween hese measu emen s and he en i onmen 23"
se e ely limi ield applica ions. Re e ence baselines a e he solu ion o unde s anding 24"
he in luence o en i onmen al condi ions. Ne e heless, he ex apola ion ou o he 25"
o iginal condi ions is no clea . The aim o his s udy was o compa e se e al 26"
app oaches o es ima e a e e ence baseline in an oli e o cha d whe e he e we e no 27"
p e ious da a om o he seasons. Two o cha ds, sepa a ed 60 m, wi h di e en ee 28"
densi y we e used. O cha d 1 had g ea e ee densi y han o cha d 2, hough he age and 29"
he cul i a we e he same. T unk diame e s o bo h o cha ds we e simila bu he c own 30"
olume o o cha d 2 was sligh ly lowe han o cha d 1. The cu en e e ence baselines 31"
o maximum daily unk diame e in bo h o cha ds we e no signi ican ly di e en 32"
be ween hem (p<0.05). In o cha d 1, he p e ious e e ence baseline was calcula ed in 33"
a 5-yea s udy ( he so called mul i-seasons app oach). The mul i-seasons app oach was 34"
no signi ican ly di e en in slope bu i was in he y-in e cep ion o he cu en 35"
e e ence baselines in bo h o cha ds (p<0.05). This app oach o e -es ima ed he alues 36"
in bo h o cha ds. Two addi ional app oaches we e es ed. These la e app oaches used 37"
da a be o e massi e pi ha dening o es ima e he cu en e e ence baseline. One o 38"
hem used he ea ly da a o es ima e a comple e e e ence baseline ( he so-called ea ly 39"
app oach). The o he ( he so-called y-ea ly app oach) used he same da a only o 40"
es ima e he y-in e cep ion and assumed ha he slope was he same as in he mul i-41"
seasons app oach. The ea ly app oach unde -es ima ed he alue o maximum daily 42"
unk sh inkage. The ea ly-y app oach p o ided a sa is ac o y es ima ion o he 43"
3"
"
e e ence baseline and imp o ed hose ob ained wi h he mul i-seasons app oach. The 44"
limi a ions and uses in i iga ion scheduling a e also discussed. 45"
46"
Keywo d: I iga ion scheduling, plan wa e s a us measu emen s, unk diame e 47"
luc ua ions, wa e ela ions. 48"
49"
50"
51"
52"
53"
54"
55"
56"
57"
58"
59"
60"
61"
62"
63"
64"
65"
66"
4"
"
1. In oduc ion 67"
The Oli e (Olea eu opaea L.) is a adi ional ain- ed ui ee in he Medi e anean 68"
basin. In he las 20 yea s, he in ensi ica ion o oli e cul i a ion has ad anced g ea ly. 69"
These new o cha ds a e i iga ed and ha e highe ee densi y han adi ional ones. The 70"
inc ease in cul i a ion has coincided wi h an inc ease in wa e demand o o he uses 71"
(Fe e es and E ans, 2006) and his s ongly limi s he amoun o wa e a ailable o 72"
i iga ion pu poses. 73"
Regula ed de ici i iga ion (RDI) is a common i iga ion scheduling echnique 74"
in ui ees, which sa es wa e wi h low o null a ia ions in yield (Behboudian and 75"
Mills, 1997; Nao , 2006). When RDI is applied, plan wa e s a us measu emen s a e 76"
needed o con ol he in ensi y o he plan wa e s ess imposed (Fe e es and González-77"
Dugo, 2009). In de ici i iga ion scheduling, echniques o con inuous measu emen o 78"
plan wa e s a us allow adequa e daily wa e ing o con ol o wa e s ess le el (O uño 79"
e al., 2010). T unk diame e luc ua ions ha e been sugges ed in se e al ui ees as a 80"
e y e icien ool o RDI (i.e. almond, Goldhame and Fe e es, 2004; peaches, 81"
Coneje o e al., 2011), bu no in o he s such as oli e ees (Mo iana e al., 2003; 82"
Mo iana e al., 2010). T unk diame e luc ua ions a e a daily cycle o sh inkage ( om 83"
he beginning o he day) and swelling ( om mid-a e noon) which occu s in all plan s 84"
(Kleppe e al., 1971). This daily cycle p o ides wo pa ame e s which a e used in 85"
i iga ion scheduling: maximum daily unk sh inkage and unk g ow h a e. 86"
Goldhame and Fe e es (2001) sugges ed he i s app oach o i iga ion 87"
scheduling wi h a unk diame e luc ua ions pa ame e . In a young o cha d, hese 88"
au ho s sugges ed he compa ison wi h maximum daily diame e o es ima e unk 89"
g ow h. Howe e , al hough his pa ame e is p esen ed in se e al s udies (i.e. 90"
Goldhame e al 1999) ex apola ion o o he loca ions is no easy. Mo iana and Fe e es 91"
5"
"
(2002), in oli e ees, epo ed ha maximum daily diame e would be di icul o 92"
ex apola e o o he condi ions and sugges ed unk g ow h a e ( he slope o maximum 93"
daily diame e ) as easie o use in i iga ion scheduling. Howe e , only in young oli e 94"
ees, a good ela ionship be ween TGR and empe a u e has been epo ed (Pé ez-95"
López e al., 2008). No s ong ela ionship be ween ei he o hese pa ame e s and a 96"
me eo ological a iable has been ound in ma u e o cha ds o any ui species. This 97"
lack o esul s is p obably ela ed o he s ong ela ionship be ween ui de elopmen 98"
and bo h pa ame e s (oli e, Mo iana e al., 2003; plum, In igliolo and Cas el, 2007; 99"
oli e, Pé ez-López e al., 2008). 100"
Maximum daily unk sh inkage is he indica o de i ed om unk diame e 101"
luc ua ions mos widely sugges ed in i iga ion scheduling in se e al ui ees (O uño 102"
e al., 2010; Fe nández and Cue as 2010). The inc ease in maximum daily unk 103"
sh inkage has adi ionally been associa ed wi h wa e s ess condi ions (O uño e al., 104"
2010), hough i is also s ongly ela ed o e apo a i e demand (He zog e al., 1995). A 105"
e e ence is he e o e needed in o de o sepa a e he e ec o e apo a i e demand and 106"
soil wa e de ici . Re e ence baseline is a simple eg ession equa ion ha es ima es 107"
maximum daily unk sh inkage in ully i iga ed condi ions om one me eo ological 108"
pa ame e . Pa ame e s such as empe a u e o apo p essu e de ici would p o ide an 109"
es ima ion o maximum daily unk sh inkage in ully i iga ed condi ions and hus 110"
i iga ion could be scheduled acco ding o he wa e s ess le el p e iously decided. 111"
Goldhame and Fe e es (2001) sugges ed he maximum daily unk diame e signal, 112"
which is he a io be ween he measu ed and he es ima ed alues, as an indica o o 113"
wa e s ess le el. 114"
P e ious s udies conduc ed on oli e ees ha e concluded ha maximum daily 115"
unk sh inkage is no a eliable wa e s a us indica o in mode a e wa e de ici 116"

6"
"
condi ions (Mo iana and Fe e es, 2002; Mo iana e al., 2010; Cue as e al., 2012). 117"
Howe e , Mo iana e al (2000) epo ed a maximum daily unk sh inkage s. s em 118"
wa e po en ial ela ionship in oli es ha es ima ed a dec ease in maximum daily unk 119"
sh inkage in se e e wa e de ici condi ions. Oli e ees a e a e y d ough esis an 120"
species, in which se e e wa e s ess condi ions du ing massi e pi ha dening (mid-121"
summe ) do no a ec o only sligh ly educe yield (Goldhame , 1999). A e e ence 122"
baseline would he e o e be use ul in o de o impose such s ess condi ions du ing his 123"
phenological pe iod. 124"
Se e al au ho s ha e epo ed e e ence baselines o maximum daily unk 125"
sh inkage in ela ion o di e en me eo ological da a. The mos commonly epo ed a e 126"
apou p essu e de ici (VPD) and ai empe a u e (amongs o he s, almond, Goldhame 127"
and Fe e es, 2004; lemon, O uño e al., 2009; manda in, Pagán e al., 2008; peach, 128"
Coneje o e al., 2011; plum, In igliolo and Cas el, 2007). The e a e only h ee s udies 129"
o oli es ha es ima ed a e e ence baseline (Mo iana and Fe e es, 2004; Mo eno e al., 130"
2006; Mo iana e al., 2011). Mo iana and Fe e es (2004), wi h one season´s da a, 131"
sugges ed VPD as he me eo ological a iable bu wi hou compa ison o o he 132"
me eo ological da a. Mo eno e al (2006), wi h one season´s da a, compa ed ou 133"
di e en me eo ological measu emen s (VPD, empe a u e, adia ion and e e ence 134"
e apo anspi a ion) and epo ed ha VPD and empe a u e p esen ed he bes 135"
ag eemen wi h maximum daily unk sh inkage da a. Mo iana e al (2011), in a i e-136"
yea s udy, showed ha maximum daily empe a u e was be eewn o he s (VPD and 137"
empe a u es a di e en ime) he bes i wi h maximum daily sh inkage. These 138"
au ho s also conside ed ha maximum empe a u e was easie o ob ain han VPD and 139"
empe a u e measu emen s a di e en ime o he day. 140"
7"
"
One o he main p oblems in he managemen o he e e ence baseline is 141"
alida ion in o he places o seasons. In o de o elimina e he in luence o in e -season 142"
changes mos s udies use da a om se e al seasons (i.e. O uño e al., 2009; Coneje o e 143"
al., 2011; Mo iana e al., 2011). Howe e , Goldhame and Fe e es (2004) epo ed a 144"
e e ence baseline es ima ed only wi h he da a o he beginning o he i iga ion season. 145"
The ex apola ion o he e e ence baseline o o he places (wi h di e en cul i a s 146"
o /and ee spaces) is mo e ela ed o ac o s such as ui load o ee dimensions and, 147"
om ou knowledge, he e a e no s udies in he li e a u e. F ui load has been epo ed 148"
as a signi ican ac o ha sligh ly a ied he equa ions ob ained (In igliolo and Cas el 149"
2007; Coneje o e al., 2010; Mo iana e al., 2011). Howe e , such di e ences a e small 150"
and, in a comme cial o cha d, ui load could be no conside ed (Mo iana e al., 2011). 151"
The in luences o ee age (Mo iana and Fe e es, 2004) o unk diame e (Gena d e al, 152"
2001; In igliolo and Cas el, 2006) we e also signi ican . Re e ence baseline would 153"
he e o e p obably need a p e ious local calib a ion. 154"
The aim o his s udy is o compa e se e al app oaches o es ima ing a e e ence 155"
baseline in an o cha d whe e he e a e no p e ious da a. This e e ence baseline will be 156"
used du ing he massi e pi ha dening pe iod. Two app oaches use cu en season da a 157"
be o e massi e pi ha dening. And a hi d app oach uses a p e ious e e ence baseline 158"
(Mo iana e al., 2011), calcula ed on he same expe imen al a m, bu in an oli e 159"
o cha d wi h close ee spacing. 160"
161"
2. Ma e ial and Me hods 162"
The expe imen was pe o med in wo o cha ds du ing he summe o 2011 a La 163"
Hampa, he expe imen al a m o he Ins i u o de Recu sos Na u ales y Ag obiología de 164"
Se illa (IRNAS-CSIC). These o cha ds a e loca ed a Co ia del Río nea Se ille (Spain) 165"
8"
"
(37º 17’’N, 6º 3’W, 30 m al i ude). The sandy loam soil (abou 2 m deep) o he 166"
expe imen al si e was cha ac e ized by a olume ic wa e con en o 0.33 m3 m-3 a 167"
sa u a ion, 0.21 m3m-3 a ield capaci y and 0.1 m3m-3 a pe manen wil ing poin , and 168"
1.30 (0-10 cm) and 1.50 (10-120 cm) g cm-3 bulk densi y. 169"
The wo oli e o cha ds (Olea eu opaea L c Manzanillo) we e i iga ed du ing 170"
he p e ious season wi h no wa e limi a ion. O cha d 1 was 41 yea s old and he ee 171"
spacing was 7 x 5 m. This o cha d was he same whe e he e e ence baseline o 172"
Mo iana e al (2011) was calcula ed, bu he ees used in he p esen s udy we e 173"
di e en . O cha d 2 was also 41 yea s old and ee spacing was 7 m x 7m. This o cha d 174"
was beside o cha d 1 and sepa a ed by a ound 60 m. The c own olume and unk 175"
diame e o he expe imen al ees we e no signi ican ly di e en be ween o cha ds 176"
(p<0.05, Table 1). Howe e , he ees in o cha d 2 had ma kedly lowe alues o c own 177"
olume han o cha d 1 (a ound 25% less) and lowe g ound co e (o cha d 1 40%; 178"
o cha d 2 24%). The beginning o he massi e pi ha dening was es ima ed acco ding o 179"
Gijón e al (2010) a day o he yea (DOY) 157 in he o cha d 1 and 164 in he o cha d 180"
2. 181"
Pes con ol, p uning and e iliza ion p ac ices we e hose commonly used by 182"
g owe s and no weeds we e allowed o de elop in he o cha d. I iga ion was ca ied ou 183"
du ing he nigh by d ip using one la e al pipe pe ee ow and i e emi e s pe plan , 184"
deli e ing 8 L h-1 each. Mic ome eo ological 30 min da a, namely ai empe a u e, 185"
sola adia ion, ela i e humidi y o ai and wind speed a 2 m abo e he soil su ace 186"
we e collec ed by an au oma ic wea he s a ion loca ed some 40 m om he 187"
expe imen al si e. Daily e e ence e apo anspi a ion (ETo) was calcula ed using he 188"
Penman-Mon ei h equa ion (Allen e al., 1998). 189"
9"
"
I iga ion equi emen s we e de e mined as he di e ence be ween c op 190"
e apo anspi a ion (ETc) and ain all. Soil mois u e was no conside ed in he wa e 191"
balance in o de o ob ain no wa e s ess condi ions. ETc was de e mined acco ding o 192"
daily e e ence e apo anspi a ion (ETo) and a c op ac o based on he ime o he yea 193"
and he pe cen age o g ound a ea shaded by he ee canopy (Fe e es and Goldhame , 194"
1990). The c op coe icien alues (Kc) conside ed we e 0.76 in May, 0.70 in June, 0.63 195"
in July and Augus , 0.72 in Sep embe and 0.77 in Oc obe (Fe nández e al., 2006). The 196"
alues o he coe icien in ela ion o he pe cen age o g ound co e ed by he c op (K ) 197"
we e 0.8 in o cha d 1 and 0.48 in o cha d 2. The alues o c op e apo anspi a ion (ETc) 198"
and o al amoun o applied wa e ( ain all no included) du ing he expe imen al pe iod 199"
( om he end o Ap il un il he middle Sep embe ) a e shown in Table 2. 200"
T unk diame e luc ua ions (TDF) a e a daily cycle o sh inkage and swelling. 201"
TDF we e measu ed h oughou he expe imen al pe iods, using a se o linea a iable 202"
displacemen ansduce s (LVDT) (model DF±2.5 mm, accu acy ±10 µm, Sola on 203"
Me ology, Bogno Regis, UK) a ached o he main unk, wi h a special b acke made 204"
o In a , an alloy o Ni and Fe wi h a he mal expansion coe icien close o ze o 205"
(Ka e ji e al., 1994). Measu emen s in 6 ees in o cha d 1 and 5 ees in o cha d 2 we e 206"
aken e e y 10 s and he da alogge (model CR10X wi h AM 416 mul iplexe , 207"
Campbell Sci. L d., Logan, USA) was p og ammed o epo 15 min means. Maximum 208"
daily unk sh inkage (MDS) was calcula ed as he di e ence be ween he maximum 209"
daily diame e , which occu s a he beginning o he day, and he minimum daily 210"
diame e , which occu s a mid-a e noon (Goldhame e al., 1999). 211"
The de e mina ion o he e e ence baseline supposed ully i iga ed condi ions. 212"
Se e al pa ame e s we e measu ed in o de o es ablish he wa e s a us o he ee (soil 213"
mois u e, maximum unk diame e , s em wa e po en ial). Soil mois u e was measu ed 214"
16"
"
es o seasons (wi h medium and high yield). The e o e, he mul i-season app oach 364"
could be use ul in condi ions o signi ican yield. 365"
Al hough signi ican yield would be expec ed an es ima ion o he e e ence 366"
baseline wi h cu en season da a and wi h he local condi ions would be a be e 367"
app oach han he mul i-season app oach. The app oach sugges ed by Goldhame and 368"
Fe e es (2004) in almonds ( he so-called ea ly app oach) was used in his way. 369"
Howe e , he ea ly app oach was he wo s in bo h o cha ds (Figs. 3 and 4). These 370"
disag eemen s wi h he esul s in almonds o Goldhame and Fe e es (2004) a e 371"
p obably ela ed o he di e ences be ween he wa e ela ions o bo h ui species. The 372"
daily cycle o s oma al lea conduc ance in ull i iga ed condi ions is no limi ed by 373"
e apo a i e demand in almond (Ma sal and Gi ona, 1997) while in oli e ees he e is a 374"
educ ion a midday (Angelopoulos e al., 1996). Such di e ences in he daily pa e n 375"
s oma a could sugges di e en espond o high anspi a ion condi ions. Acco ding o 376"
he p esen s udy, maximum daily unk sh inkage g ea e han 400 µm was unde -377"
es ima ed wi h he ea ly app oach (Fig. 4). The e o e, he g ea es condi ions o 378"
anspi a ion likely changed he amoun o wa e ha he unk ans e ed o he 379"
anspi a ion s eam. In oli e, he ee could inc eases he amoun o wa e om he 380"
unk mo e han in almond and hen maximum daily sh inkage was highe han 381"
expec ed, Such inc ease could be ela ed o he g ea e capaci y o dehyd a ion in oli e 382"
ees (Fe e es, 1984). 383"
Maximum daily unk sh inkage is a ec ed by se e al ac o s which would 384"
change he pa e n o i s ela ionship wi h empe a u e in he long e m, wa e s a us 385"
condi ions (Gena d e al., 2001) and wood composi ion (D ew and Dones, 2009) a e 386"
p obably he mos impo an . Such changes would p obably a y he e e ence baseline 387"
e en hough ully-i iga ed condi ions we e pe o med du ing he cu en o p e ious 388"

17"
"
season. The p esen s udy con i med ha hese changes occu ed, bu only in he y-389"
in e cep ion, while he slope o he equa ion was no a ec ed (Fig. 4). The ea ly-y 390"
app oach made i possible o ob ain he e e ence baseline be o e massi e pi ha dening 391"
wi h only he es ima ion o he y-in e cep ion. The ex apola ion o o he Manzanillo 392"
o cha ds is p obably possible, bu u he wo k is needed o s udy he in luence o he 393"
cul i a . Mo iana and Fe e es (2004) epo ed e e ence baseline using apo p essu e 394"
de ici (VPD) wi h di e en cul i a s and age o cha ds, bu simila en i onmen al 395"
condi ions. When VPD is conside ed o he same ui load an almos equal slope was 396"
ound be ween ma u e c Manzanillo (Mo iana e al. 2011) and ma u e c Picual 397"
(Mo iana and Fe e es, 2004). The in luence o ee age is p obably di e en , young c 398"
A bequino showed a di e en slope and y-in e cep ion han ma u e c Picual (Mo iana 399"
and Fe e es, 2004) and ma u e c Manzanillo (Mo iana e al., 2011). Goldhame and 400"
Fe e es (2001) sugges ed ha he maximum daily unk sh inkage would be smalle in 401"
young ees han in ma u e because o he g ea e g ow h in he o me . In almonds, he 402"
e e ence baseline epo ed in se e al s udies o young o cha ds was simila wi h 403"
di e en cul i a s (Fe e es and Goldhame , 2003; Goldhame and Fe e es, 2004). Egea 404"
e al. (2009) wi h h ee season´s da a in a young almond o cha d sugges ed ha he 405"
e e ence baseline could be es ima ed e e y 1-2 yea s wi h he da a o he beginning o 406"
he season, hough sligh di e ences due o phenological s age we e epo ed. 407"
408"
5. Conclusions 409"
The ex apola ion o e e ence baseline o maximum daily unk sh inkage o o he 410"
o cha ds was possible in ma u e cul i a Manzanillo. The e e ence baselines we e 411"
simila be ween o cha ds, hough spacing and g ound co e we e di e en . Sligh 412"
a ia ions in he y-in e cep ion we e ound in bo h e e ence equa ions. The mul i-413"
18"
"
season app oach, a gene al equa ion p e iously calcula ed, o e -es ima ed bo h 414"
e e ence baselines. Such esul s we e p obably ela ed o he ex emely low ui load in 415"
bo h o cha ds in compa ison wi h he ones ob ained when he mul i-season app oach 416"
was calcula ed. A mul i-season app oach would be use ul in condi ions o signi ican 417"
yield. An ea ly app oach ha es ima ed he comple e (slope and y-in e cep ion) 418"
e e ence baseline was no possible because signi ican a ia ions in he slope we e 419"
ound wi h he inc ease in maximum daily unk sh inkage. This esul is no consis en 420"
wi h he ones epo ed in almond. The di e ences in he s oma a lea conduc ance 421"
pa e n be ween bo h species we e p obably ela ed o he lack o esul s. The e e ence 422"
baseline in all he es ima ions was simila in slope bu di e en in y-in e cep ion. The 423"
ea ly-y app oach ha only es ima ed his la e componen o he e e ence baseline 424"
p esen ed a good i be ween obse ed and measu ed da a du ing he pe iod o pi 425"
ha dening. Acco ding o he p esen esul s and li e a u e he ea ly-y app oach would be 426"
use ul in he es ima ion o e e ence baseline in ma u e o cha ds e en when di e en 427"
cul i a s we e conside ed. 428"
429"
Acknowledgemen s 430"
This esea ch was suppo ed by he Spanish Minis e io de Economia y Compe i i idad 431"
(MINECO), (AGL2010-19201-CO4-03 p ojec ). The s ay o D. Mo ales and J. 432"
Dell’Amico was suppo ed by he AECID p ojec D/030431/10. Thanks a e due o A. 433"
Mon e o and J. Rod iguez o help wi h ield measu emen s and o p o esso s M. Ce a 434"
and C. Co és o help in he s a is ical analysis. 435"
436"
437"
19"
"
Re e ence 438"
Allen, R.G., Pe ei a, L.S., Raes, D., Smi h, M. 1998. C op e apo anspi a ion. 439"
Guideline o compu ing c op wa e equi emen s. FAO i iga ion and d ainage 440"
pape nº 56. Roma.FAO. 441"
Angelopoulos, K., Dichio, B., Xiloyannis, C. 1996. Inhibi ion o pho osyn hesis in oli e 442"
ees (Olea eu opaea L.) du ing wa e s ess and ewa e ing. J Exp Bo 47, 1093-443"
110. 444"
Behboudian, M.H., Mills, T.M., 1997. De ici i iga ion in deciduous o cha ds. Ho . 445"
Re . 21, 105-131. 446"
B ough, D.W., Jones, H.G., G ace, J. 1986. Diu nal changes in wa e con en o he 447"
s ems o apple ees, as in luenced by i iga ion. Plan Cell En i on. 9, 1-7. 448"
Coneje o, W., Mellisho, C.D., O uño, M.F., Galindo, A., Pé ez-Sa mien o, F., 449"
To ecillas, A. 2010. Es ablishing máximum daily unk sh inkage and midday 450"
s em wá e po en ial e e ence equa ions o i iga ion scheduling o ea ly 451"
ma u ing peach ees. I ig. Sci. 29, 299-309. 452"
Coneje o, W., Mellisho, C. D., O uno, M. F., Mo iana, A., Mo eno, F., To ecillas, A. 453"
2011. Using unk diame e senso s o egula ed de ici i iga ion scheduling in 454"
ea ly ma u ing peach ees. En i on. Exp. Bo . 71, 409-415. 455"
Cue as MV, Ma ín-Palomo MJ, Díaz-Espejo A, To es-Ruíz JM, Rod íguez-456"
Dominguez CM, Pé ez-Ma ín A, Pino-Mejías R, Fe nández JE (2012). 457"
Assessing wa e s ess in a hedge ow oli e o cha d om sap lo and unk 458"
diame e measu emen s. I ig. Sci. doi 10.1007/s00271-0.12-0357-x. 459"
D ew, D.M., Downes, G.M. 2009. The use o p ecision dend ome e s in ease ch on 460"
daily s em size and wood p ope y a ia ion: A e iew. Dend och onologia 27, 461"
159-172. 462"
20"
"
Fe e es, E. 1984. Va iabili y in adap i e mechanisms o wa e de ici s in annual and 463"
pe ennial c op plan s. Bull. Soc. Bo . F ance. 131, 17-32. 464"
Fe e es, E., E ans, R.G. 2006. I iga ion o ui ees and ines: an in oduc ion. I ig. 465"
Sci. 24, 55-57. 466"
Fe e es, E., Goldhame , D. 1990. Deciduous ui and nu ees. In: S ewa , B.A., 467"
Nielsen D.R. (Eds), I iga ion o Ag icul u al C ops. Ag onomy Monog aph No. 468"
30. Ame ican Socie y o Ag onomy, Madison, pp. 987-1017. 469"
Fe e es, E., Goldhame , D. 2003. Sui abili y o s em diame e a ia ions and wa e 470"
po en ial as indica o s o i iga ion scheduling o almond ees. J. Ho . Sci. 471"
Bio ech. 78,139-144. 472"
Fe e es, E., González-Dugo, V. 2009. Imp o ing p oduc i i y o ace wa e sca ci y in 473"
i iga ed ag icul u e. In: C op Physiology: applica ions o gene ic imp o emen s 474"
and ag onomy. Sad as, V. and Calde ini, D. (eds). Pp 122-143. Ams e dam, 475"
Academic P ess. 476"
Fe nández, J. E.; Cue as, M. V. 2010. I iga ion scheduling om s em diame e 477"
a ia ions: A e iew. Ag . Fo es Me eo ol., 150, 135-151. 478"
Gena d, M., Fishman, S., Ve camb e, G., Hugue , J.G., Bussi, C., Besse , J., Habib, R. 479"
2001. A biophysical analysis o s em and oo diame e a ia ions in woody 480"
plan s. Plan Physiol. 126, 188-202. 481"
Gijón, M.C., Pé ez-López, D., Gue e o, J., Coucei o, J.F., Mo iana, A. 2010. Riego 482"
de ici a io con olado en oli o y pis ache o. Ag icul u a 930, 458-462 483"
Goldhame , D.A. 1999. Regula ed de ici i iga ion o Cali o nia canning oli es. Ac a 484"
Ho ic. 474, 369-372. 485"
21"
"
Goldhame D.A.; Fe e es E. 2001. I iga ion scheduling p o ocols using con inuously 486"
eco ded unk diame e measu emen s. I ig. Sci. 20, 115-125 487"
Goldhame , DA; Fe e es, E 2004. I iga ion scheduling o almond ees wi h unk 488"
diame e senso s. I iga ion Sci. , 23: 11-19. 489"
Goldhame , D., Fe e es, E., Ma a, M., Gi ona, J., Cohen, M. 1999. Sensi i i y o 490"
con inuous and disc e e plan and soil wá e s a us moni o ing in peach ees 491"
subjec ed o de ici i iga ion. J. Am. Soc. Ho ic. Sci. 124, 437-444. 492"
He zog K.M., Hüsle R., Thum R. 1995. Diu nal changes in he adius o a subalpine 493"
No way sp uce s em: hei ela ion o he sap low and hei use o es ima e 494"
anspi a ion. T ees 10, 94-101. 495"
In igliolo, D.S., Cas el, J.R. 2006. Use ulness o diu nal unk sh inkage as a wa e 496"
s ess indica o in plum ees. T ee Physiol., 26 303–311 497"
In igliolo, D.S., Cas el, J.R. 2007. C op load a ec s maximum daily unk sh inkage o 498"
plum ees. T ee Physiol., 27: 89-96. 499"
Ka e ji,N., Ta dieu,F.,Be henod,O., Que in, P. 1994. Beha io o Maize s em diame e 500"
du ing d ying cycles:compa ison o wo me hods o de ec ing wa e s ess. C op 501"
Sci. 34, 165-169. 502"
Kleppe , B., B owing, V.D., Taylo , H.M. 1971. S em diame e in ela ion o plan wa e 503"
s a us. Plan Physiol. 48, 683-685. 504"
Ma sal, J., Gi ona, J. 1997. Lea wa e ela ions pa ame e s in almond compa ed o 505"
hazelnu ees du ing a de ici i iga ion pe iod. J. Ame . Soc. Ho . Sci. 122, 506"
582-587. 507"
Ma ín-Ve edo , A.I., Pé ez-Rod íguez, J.M., P ie o-Losada, H., Fe e es, E. 2011. 508"
In e ac i e esponses o wa e de ici s and c op load in oli e (Olea eu opaea L., 509"
c . Mo isca) I. - G ow h and wa e ela ions. Ag . Wa e Manage. 98, 941-949. 510"

22"
"
Mo eno, F., Coneje o, W., Ma in-Palomo, M. J., Gi on, I. F., To ecillas, A., 2006. 511"
Maximum daily unk sh inkage e e ence alues o i iga ion scheduling in 512"
oli e ees. Ag . Wa e Manage. 84, 290-294. 513"
Mo iana, A., Fe e es, E. 2002. Plan indica o s o scheduling i iga ion o young oli e 514"
ees. I ig. Sci. 21, 83-90. 515"
Mo iana, A., Fe e es, E. 2004. Es ablishing e e ences alues o unk diame e 516"
luc ua ions and s em wa e po en ial o i iga ion scheduling o oli e ees. 517"
Ac a Ho ic. 664, 407-412. 518"
Mo iana, A., Fe e es, E., O gaz, F., Cas o, J., Humanes, M.D., Pas o , M. 2000. The 519"
ela ions be ween unk diame e luc ua ions and ee wa e s a us in oli e ee 520"
(Oleaeu opea L). Ac a Ho ic. 537, 293-297. 521"
Mo iana, A., O gaz, F., Fe e es, E., Pas o , M. 2003. Yield esponses o a ma u e oli e 522"
o cha d o wa e de ici s. J. Am. Soc. Ho ic. Sci..128, 425-431. 523"
Mo iana, A., Gi on, I.F., Ma in-Palomo, M.J., Coneje o, W., O uno, M.F., To ecillas, 524"
A., Mo eno, F. 2010. New app oach o oli e ees i iga ion scheduling using 525"
unk diame e senso s. Ag ic. Wa e Manage. 97, 1822-1828. 526"
Mo iana, A., Mo eno, F., Gi on, I.F., Coneje o, W. O uño, M.F., Mo ales, D., Co ell, 527"
M., To ecillas, A. 2011. Seasonal changes o maximum daily sh inkage 528"
e e ence equa ions o i iga ion scheduling in oli e ees: In luence o ui 529"
load. Ag ic. Wa e Manage. 99, 121-127. 530"
Nao , A. 2006. I iga ion scheduling and e alua ion o ee wa e s a us in deciduous 531"
o cha d. Ho . Re . 32, 111-165. 532"
Nao , A., Gal. Y., Pe es, M. 2006. The inhe en a iabili y o wa e s ess indica o s in 533"
apple, nec a ine and pea o cha ds and he alidi y o a lea selec ion p ocedu e 534"
o wa e po en ial measu emen s. I ig. Sci. 24,129-135. 535"
23"
"
O uño, M. F., Ga cia-O ellana, Y., Coneje o, W. 2009. Assessmen o maximum daily 536"
unk sh inkage signal in ensi y h eshold alues o de ici i iga ion in lemon 537"
ees. Ag ic. Wa e Manage. 96, 80-86. 538"
O uño, M.F., Coneje o, W., Mo eno, F., Mo iana, A., In igliolo, D.S., Biel, C., 539"
Mellisho, C.D., Pé ez-Pas o , A., Domingo, R., Ruiz-Sánchez, M.C., Casadesus, 540"
J., Bonany, J., To ecillas, A. 2010. Could unk diame e senso s be used in 541"
woody c ops o i iga ion scheduling? A e iew o cu en knowledge and 542"
u u e pe spec i es. Ag ic. Wa e Manage. 97, 1–11. 543"
Pagán, E., Pé ez-Pas o , A., Domingo, R., Conesa, M.R., Robles, J.M., Bo ía, P., 544"
Ga cía-Olle , I., Ca o, M. 2008. Feasibili y s udy o he maximum daily unk 545"
sh inkage o scheduling manda in ees i iga ion. I al. J. Ag on. 3, 691-692. 546"
Pé ez-López, D., Mo iana, A., Rapopo , H., Olmedilla, N., Ribas, F. 2008. New 547"
app oach o using unk g ow h a e and endoca p de elopmen in he i iga ion 548"
scheduling o young oli e o cha d. Sci. Ho . 115, 244-251. 549"
Scholande , P.F., Hammel, H.T., B ads ees , E.A., Hemmingsen, E.A. 1965. Sap 550"
p essu e in ascula plan . Sci. 148, 339-346. 551"
Simonneau, T., Habib, R., Gou ouly, J.P., Hugue , J.G. 1993. Diu nal changes in s em 552"
diame e depend upon a ia ions in wa e con en : di ec e idence in peach ees. 553"
J. Exp. Bo . 44, 615-621. 554"
555"
556"
557"
558"
559"
24"
"
560"
Table 1. Dimensions and dis ance in he o cha d o he expe imen . The e we e no 561"
signi ican di e ences be ween o cha d 1 and o cha d 2 in c own olume o unk 562"
diame e . 563"
564"
565"
566"
567"
568"
569"
570"
571"
572"
573"
574"
575"
576"
Dis ance
(mxm)
G ound co e
C own Volume
(m3 ee-1)
T unk
Diame e
(m)
O cha d 1
7x5
40%
31.3±4
0.24±0.01
O cha d 2
7x7
24%
23.6±2
0.23±0.01
25"
"
577"
Table 2. C op e apo anspi a ion, i iga ion applied and yield componen s: yield (Yield
(kg ee-1) and ha es ( ha-1).
ETc
(mm)
I iga ion
applied
(mm)
Yield
(kg ee-1)
Yield
( ha-1)
O cha d 1
299.7
285.4
8.9 ± 6.0
2.5 ± 1.7
O cha d 2
214.0
196.7
2.9 ± 1.4
0.6 ± 0.3
578"
579"