Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae)

0864 Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae) André Larochelle and Marie-Claude Larivière New Zealand Arthropod Collection, Manaaki Whenua–Landcare Research, Private Bag 92170, Auckland 1142, New Zealand Center for Systematic Entomology, Inc., Gainesville, FL Date of issue: April 30, 2021 Insecta MundI A journal of world insect systematics Page Count: 96 Larochelle and Larivière Prosphodrus waimana Larochelle and Larivière, new species Larochelle A, Larivière M-C. 2021. Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae). Insecta Mundi 0864: 1–96. Published on April 30, 2021 by Center for Systematic Entomology, Inc. P.O. Box 141874 Gainesville, FL 32614-1874 USA http://centerforsystematicentomology.org/ Insecta Mundi is a journal primarily devoted to insect systematics, but articles can be published on any nonmarine arthropod. 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This is an open access article distributed under the terms of the Creative Commons, Attribution Non-Commercial License, which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original author(s) and source are credited. http://creativecommons.org/licenses/by-nc/3.0/ Insecta MundI0864: 1–96 2021 Synopsis of the tribe Platynini in New Zealand (Coleoptera: Carabidae) André Larochelle New Zealand Arthropod Collection, Manaaki Whenua–Landcare Research Private Bag 92170, Auckland 1142, New Zealand [email protected] Marie-Claude Larivière New Zealand Arthropod Collection, Manaaki Whenua–Landcare Research Private Bag 92170, Auckland 1142, New Zealand [email protected] Abstract. The tribe Platynini (Coleoptera: Carabidae: Harpalinae) is revised for New Zealand. Eight genera and forty-three species are recognized. Four genera and sixteen species are described as new: Ctenognathus davidsoni Larochelle and Larivière new species, Ctenognathus earlyi Larochelle and Larivière new species, Ctenognathus garnerae Larochelle and Larivière new species, Ctenognathus hoarei Larochelle and Larivière new species, Ctenognathus kaikoura Larochelle and Larivière new species, Ctenognathus marieclaudiae Larochelle new species, Ctenognathus perumalae Larochelle and Larivière new species, Ctenognathus takahe Larochelle and Larivière new species, Ctenognathus tawanui Larochelle and Larivière new species, Ctenognathus tepaki Larochelle and Larivière new species, Ctenognathus urewera Larochelle and Larivière new species, Kiwiplatynus Larochelle and Larivière new genus, Kiwiplatynus taranaki Larochelle and Larivière new species, Kupeplatynus Larochelle and Larivière new genus, Maoriplatynus Larochelle and Larivière new genus, Maoriplatynus marrisi Larochelle and Larivière new species, Prosphodrus mangamuka Larochelle and Larivière new species, Prosphodrus sirvidi Larochelle and Larivière new species, Prosphodrus waimana Larochelle and Larivière new species, Tuiplatynus Larochelle and Larivière new genus. Lectotypes are designated for twelve taxa: Anchomenus adamsi Broun, 1886, Anchomenus colensonis White, 1846, Anchomenus feredayi Bates, 1874, Anchomenus helmsi Sharp, 1881, Anchomenus intermedius Broun, 1908, Anchomenus macrocoelis Broun, 1908, Anchomenus munroi Broun, 1893, Anchomenus sophronitis Broun, 1908, Colpodes crenatus Chaudoir, 1878, Colpodes neozelandicus Chaudoir, 1878, Ctenognathus littorellus Broun, 1908, and Ctenognathus pictonensis Sharp, 1886. Six new combinations are established: Ctenognathus bidens (Chaudoir, 1878) = Kiwiplatynus bidens (Chaudoir, 1878); Ctenognathus crenatus (Chaudoir, 1878) = Kupeplatynus crenatus (Chaudoir, 1878); Ctenognathus lucifugus (Broun, 1886) = Kupeplatynus lucifugus (Broun, 1886); Ctenognathus sulcitarsis (Broun, 1880) = Kupeplatynus sulcitarsis (Broun, 1880); Ctenognathus libitus (Broun, 1914) = Tuiplatynus libitus (Broun, 1914); Ctenognathus sophronitis (Broun, 1908) = Tuiplatynus sophronitis (Broun, 1908). Fifteen new synonymies are established: Ctenognathus littorellus Broun, 1908 = Ctenognathus adamsi (Broun, 1886); Anchomenus parabilis Broun, 1880 = Ctenognathus cardiophorus (Chaudoir, 1878); Anchomenus integratus Broun, 1908 = Ctenognathus colensonis (White, 1846); Anchomenus macrocoelis Broun, 1908 = Ctenognathus edwardsii (Bates, 1874); Ctenognathus actochares Broun, 1894 = Ctenognathus elevatus (White, 1846); Anchomenus punctulatus Broun, 1877, Anchomenus montivagus Broun, 1880, Anchomenus perrugithorax Broun, 1880, Anchomenus politulus Broun, 1880, Anchomenus suborbithorax Broun, 1880, and Colpodes neozelandicus Chaudoir, 1878 = Ctenognathus novaezelandiae (Fairmaire, 1843); Ctenognathus simmondsi Broun, 1912 = Ctenognathus pictonensis Sharp, 1886; Anchomenus (Platynus) cheesemani Broun, 1880 and Calathus deformipes Broun, 1880 = Kupeplatynus crenatus (Chaudoir, 1878); Anchomenus munroi Broun, 1893 = Kupeplatynus lucifugus (Broun, 1886). Ctenognathus elevatus (White, 1846), previously synonymized with Ctenognathus novaezelandiae (Fairmaire, 1843), is reinstated as full species. A revision of all taxa is provided. Descriptions, identification keys, illustrations of male genitalia, habitus photos, distributional data and maps are given. Extensive information on ecology, biology, dispersal power, and collecting techniques is included for each species. Key words. Taxonomy, new genera and species, keys, geographic distribution, ecology, biology, dispersal power. ZooBank registration. urn:lsid:zoobank.org:pub:DF505A18-63A1-44BB-BF5D-13887FAE0DAD 2 · April 30, 2021 Larochelle and Larivière Introduction The tribe Platynini (Carabidae: Harpalinae) is worldwide in distribution. Members of this group occur throughout New Zealand, except on subantarctic islands. They are mostly forest-dwellers and are especially abundant in the vicinity of streams. The New Zealand Platynini were catalogued by Larochelle and Larivière (2001) in eight genera: “Anchomenus” Bonelli, Cerabilia Laporte de Castelnau, Ctenognathus Fairmaire, Laemostenus Bonelli, Notagonum Darlington, Platynus Bonelli, Prosphodrus Britton, and Zabronothus Broun. New Zealand species previously recorded in the genus “Anchomenus” were transferred to the genus Ctenognathus by Liebherr (2005) and Larochelle and Larivière (2007). The genus Laemostenus belongs to the tribe Sphodrini considered separate from the tribe Platynini by Lorenz (2005), Bouchard et al. (2011), and Bousquet (2012). Larochelle and Larivière (2016) adopted this position in their Taxonomic Supplement to the 2001 New Zealand catalogue. One species of Notagonum, N. marginellum (Erichson, 1842), was deleted from the New Zealand fauna by Larochelle and Larivière (2007); this species occurs in Tasmania, southern mainland Australia, and on Lord Howe Island (Baehr 2016). The genus Zabronothus was synonymized with Cerabilia by Larochelle and Larivière (2007). Subsequently, Cerabilia was transferred from Platynini to Loxandrini (Will 2011) and then to Abacetini (Will 2015). These taxonomic changes resulted in four genera (Ctenognathus, Notagonum, Platynus, Prosphodrus) and 41 species of Platynini being recorded from New Zealand in Larochelle and Larivière (2016). Between 1877 and 1921, Broun described the bulk of the platynines for New Zealand. Since then, only two species have been described for this country, Prosphodrus waltoni Britton, 1959 and P. occultus Britton, 1960. The current taxonomic revision adds four genera to the fauna (Kiwiplatynus new genus, Kupeplatynus new genus, Maoriplatynus new genus, Tuiplatynus new genus) and deals with 43 endemic species, 16 of which are new to science. This synopsis provides a detailed treatment of the taxonomy of New Zealand Platynini, identification keys to all taxa, and extensive information on species distribution, ecology, biology, dispersal power, and collecting techniques. The male genitalia are described and illustrated, and habitus photos are provided for all taxa for the first time. This revision is another step in the authors’ goal of attaining a comprehensive understanding of the New Zealand carabid fauna within a reasonable time frame, and of making large amounts of information available for practical use by a wide range of users. It follows the publication of a catalogue of New Zealand Carabidae (Larochelle and Larivière 2001), a revision of the tribe Harpalini (Larochelle and Larivière 2005), a synopsis of supraspecific carabid taxa (Larochelle and Larivière 2007), a synopsis of species of the tribes Amarotypini, Cicindelini, Clivinini, Migadopini, Pamborini, Rhysodini, Moriomorphini, and Trechini (Larochelle and Larivière 2013), a synopsis of the genus Bembidion Latreille (Larochelle and Larivière 2015), a taxonomic supplement (2001 to 2015) to the 2001 catalogue (Larochelle and Larivière 2016), and a synopsis of the tribe Zolini (Larochelle and Larivière 2017). Materials and Methods This study is based on the examination of about 15,000 specimens from several hundred New Zealand localities. Most of this material (about 70%) was collected by the authors from 1992 to 2018 and is deposited in the New Zealand Arthropod Collection (NZAC), Auckland. Other specimens were kindly provided by the following museums and collections: Auckland War Memorial Museum, Auckland, New Zealand (AMNZ); Canterbury Museum, Christchurch, New Zealand (CMNZ); Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, U.S.A. (CMNH); Entomology Research Collection, Lincoln University, Lincoln, New Zealand (LUNZ); John Nunn private collection, Dunedin, New Zealand (JNNZ); Muséum national d’Histoire naturelle, Paris, France (MNHN); Museum of New Zealand Te Papa Tongarewa, Wellington, New Zealand (MONZ); The Natural History Museum, London, U.K. (BMNH). Type material has been deposited in some of the above collections. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 3 The morphological terminology used in this work generally follows Larochelle and Larivière (2007, 2013, 2015, 2017). All descriptions are based on the same list of characters so as to be fully comparative between taxa. The microsculpture (Fig. 1–4) of the head, pronotum, and elytra, the mentum tooth (Fig. 14–18), and the metatarsomeres 4 (Fig. 29–31), were examined in great detail and proved highly useful in discriminating taxa. The male genitalia, also highly diagnostic at the species level, were dissected across numerous populations of each taxon. In the identification keys to genera and species, additionally helpful but not necessarily exclusive characters are provided in brackets. The taxonomic arrangement of genera and species, and the sequence of habitus photos and illustrations of male genitalia, follow the order of taxa in the identification keys. Synonyms, new combinations, and type data are given for all taxa. Type data are listed in this order: type status followed by sex, acronym of entomological collection or museum serving as repository, and original label data with a forward slash (/) indicating a different label. The two-letter abbreviation codes of Crosby et al. (1976, 1998) for areas of New Zealand (Fig. 123) were used to record geographic distributions. Full distributional information is given for species known from 10 or fewer localities. Appendix 1 provides decimal degree coordinates for localities cited in the text. Maps summarizing species distribution are alphabetically arranged (Fig. 124–166). Notes on the ecology, biology, and dispersal power are based on an analysis and synthesis of specimen label data and field observations by the authors. The terminology and style of presentation follow Larochelle and Larivière (2001, 2003). Habitus photos were taken by B. E. Rhode. The authors prepared all other illustrations. Tribe Platynini Description (New Zealand). Body length 7.0–26.2 mm; not pedunculate. Head. Mandibles with setiferous puncture in scrobe. Labrum subtruncate or emarginate anteriorly. Eyes present, touching buccal fissures or separated from them; two setiferous punctures on inner side of each eye (a single posterior puncture in Kupeplatynus). Tempora inflated or not. Clypeus with a setiferous puncture on each side. Antennae filiform; segments 1–3 with a few setae, segments 4–11 densely pubescent; segments 3–10 with verticillate setae. Mentum: medial tooth present; two circular foveae; usually two long setae (two, four or six long setae in Prosphodrus; 10–18 short setae in addition to the two long setae in Tuiplatynus). Mentum-submentum suture present. Submentum with two to twelve long setae (twelve short setae in addition to two long setae in Maoriplatynus). Ligula with two setae. Paraglossae glabrous (pubescent in Prosphodrus). Palpi: terminal segment obtuse or truncate apically; terminal and penultimate maxillary segments glabrous; penultimate labial segment with two setae on anterior margin. Thorax. Pronotum with a single setiferous puncture (anteriorly) or with two setiferous punctures on each side (anteriorly and posteriorly). Scutellum visible, inserted entirely between elytral bases. Legs. Metacoxae with two posteroventral setae (three or four setae in Kiwiplatynus and Kupeplatynus). Protibiae without outer apical prolongation. Tarsi glabrous dorsally; basal segments 1–3 of male protarsi dilated, with two rows of ventral scales; metatarsomeres 4 bilobed apically; claws entire ventrally; unguitractor plate invisible between tarsal claws. Elytra. Usually fused along suture (hindwings vestigial, or, in some Notagonum, about half developed), or free along suture (hindwings fully developed, in some Notagonum and in Platynus). Basal margin usually complete, reaching about scutellum (incomplete in Kupeplatynus (in part) and Prosphodrus (in part)). Interval 3 without setiferous punctures or with one to three setiferous punctures. Umbilicate series with 13–25 setiferous punctures. Radial field without fine dense pubescence. Apices usually obtuse or rounded, rarely mucronate or truncate-emarginate (turned upward in Maoriplatynus). Epipleura simple (without inner fold or plica) near apex. Abdomen. Apex invisible dorsally. Sterna IV–VI: both sexes usually with two long apical ambulatory setae (additional long ambulatory setae in Kiwiplatynus). Sternum VII: male with two or four long apical ambulatory setae; female usually with four, eight, or ten long apical ambulatory setae (rarely with two or six long apical ambulatory setae). Sterna V–VII: both sexes with additional numerous short setae in Maoriplatynus. Aedeagus. Lateral view: moderately or strongly arcuate (slightly arcuate in Maoriplatynus); basal lobe of basal bulb usually present (absent in Maoriplatynus and Ctenognathus (in part)); internal sac unarmed (armed with scale-like sclerites in Kiwiplatynus and Kupeplatynus (in part)). Dorsal view: narrow to very wide, asymmetrical (ostium of membranous area deflected to the left) or symmetrical (ostium dorsal); basal bulb moderately distant from membranous area or close to it. Parameres: left paramere larger, oval; right paramere slender, often shorter. 4 · April 30, 2021 Larochelle and Larivière References. Darlington 1952: 89–252 (revision, New Guinean Platynini), 1956: 1–10 (key, Australian genera); Lindroth 1966: 552–648, 1968: 649–650 (revision, North American Platynini); Habu 1978: 1–447 (revision, Japanese Platynini); Liebherr and Zimmerman 1998: 137–172 (cladistics analysis, phylogeny, biography, Hawaiian Platynini), 2000: 1–494 (revision, Hawaiian Platynini); Larochelle and Larivière 2001: 128–140 (catalogue); Liebherr 2005: 263–295 (revision, Vanuatu Platynini); Larochelle and Larivière 2007: 78–82 (taxonomy), 2016: 36–38 (list). Alphabetical checklist of taxa Valid genusand species-group taxa are listed alphabetically (A = Adventive, E = Endemic, N = Native, but not endemic to New Zealand). Tribe Platynini Genus Ctenognathus Fairmaire, 1843 E adamsi (Broun, 1886) E arnaudensis (Broun, 1921) E cardiophorus (Chaudoir, 1878) E colensonis (White, 1846) E davidsoni Larochelle and Larivière E, new species earlyi Larochelle and Larivière E, new species edwardsii (Bates, 1874) E elevatus (White, 1846) E garnerae Larochelle and Larivière E, new species helmsi (Sharp, 1881) E hoarei Larochelle and Larivière E, new species intermedius (Broun, 1908) E kaikoura Larochelle and Larivière E, new species marieclaudiae Larochelle E, new species novaezelandiae (Fairmaire, 1843) E oreobius (Broun, 1886) E otagoensis (Bates, 1878) E perumalae Larochelle and Larivière E, new species pictonensis Sharp, 1886 E sandageri (Broun, 1882) E takahe Larochelle and Larivière E, new species tawanui Larochelle and Larivière E, new species tepaki Larochelle and Larivière E, new species urewera Larochelle and Larivière E, new species xanthomelus (Broun, 1908) E Genus Kiwiplatynus Larochelle and Larivière E, new genus bidens (Chaudoir, 1878) E taranaki Larochelle and Larivière E, new species Genus Kupeplatynus Larochelle and Larivière E, new genus crenatus (Chaudoir, 1878) E lucifugus (Broun, 1886) E sulcitarsis (Broun, 1880) E Genus Maoriplatynus Larochelle and Larivière E, new genus marrisi Larochelle and Larivière E, new species Genus Notagonum Darlington, 1952 N chathamense (Broun, 1909) E feredayi (Bates, 1874) E lawsoni (Bates, 1874) E submetallicum (White, 1846) E Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 5 Genus Platynus Bonelli, 1810 N macropterus (Chaudoir, 1879) E Genus Prosphodrus Britton, 1959 E mangamuka Larochelle and Larivière E, new species occultus Britton, 1960 E sirvidi Larochelle and Larivière E, new species waimana Larochelle and Larivière E, new species waltoni Britton, 1959 E Genus Tuiplatynus Larochelle and Larivière E, new genus libitus (Broun, 1914) E sophronitis (Broun, 1908) E Key to genera of Platynini (New Zealand) 1. Eyes (Fig. 8) touching buccal fissures; hindwings fully developed or half developed; scutellar striole of elytra very long ...................................................................... 2 — Eyes (Fig. 9) separated from buccal fissures; hindwings vestigial; scutellar striole of elytra short or moderately long .......................................................................... 3 2(1). Mesoand metatarsomeres 1–4 (Fig. 28) tricarinate dorsally; forebody narrow in comparison to elytra; body without metallic luster. [Larger, body length 13.0–14.2 mm; North Island, South Island, and Stewart Island] ...................................................Genus Platynus Bonelli — Mesoand metatarsomeres 1–4 (Fig. 27) not carinate dorsally; forebody moderately wide in comparison to elytra; body with metallic luster. [Smaller, body length 7.3–10.4 mm; North Island, South Island, Stewart Island, and Offshore Islands (TH, CH)] ................Genus Notagonum Darlington 3(1). Paraglossae (Fig. 12) pubescent; prosternal lobe compressed into a vertical ridge. [Body length 14.0–26.2 mm; mentum tooth bifid; abdominal sterna V–VII without short setae; North Island] ........... .............................................................Genus Prosphodrus Britton — Paraglossae (Fig. 13) glabrous; prosternal lobe not compressed into a vertical ridge ............... 4 4(3). Abdominal sterna V–VII (Fig. 34) with numerous short setae in addition to long ambulatory setae; elytral apices turned upward; mentum tooth bifid (Fig. 14); aedeagus (in lateral view) slightly arcuate (Fig. 90). [Body length 12.5–12.7 mm; Three Kings Islands (TH)] ............................. ................................ Genus Maoriplatynus Larochelle and Larivière, new genus — Abdominal sterna V–VII without short setae; elytral apices not turned upward; mentum tooth entire, subtruncate (Fig. 16), acute (Fig. 18), or emarginate (Fig. 15); aedeagus (in lateral view) more strongly arcuate ...................................................................... 5 5(4). Cleaning organ of protibiae (Fig. 26) with three clip setae; metacoxae with three or four posteroventral setae (Fig. 23); neck constriction deep dorsolaterally (Fig. 5) ............................... 6 — Cleaning organ of protibiae (Fig. 25) with two clip setae; metacoxae with two posteroventral setae (Fig. 24); neck constriction shallow (Fig. 6) or absent (Fig. 7) dorsolaterally ....................... 7 6(5). Abdominal sterna V–VI (Fig. 32) of both sexes with three to five long apical ambulatory setae; two setiferous punctures on each side of each eye; forebody narrow in comparison to elytra; interval 3 of elytra with one or two setiferous punctures; mentum tooth (Fig. 15) emarginate apically. [Body length 16.0–19.3 mm; northern North Island] .............................................. .................................. Genus Kiwiplatynus Larochelle and Larivière, new genus — Abdominal sterna V–VI (Fig. 33) of both sexes with two long apical ambulatory setae; a single setiferous puncture on each side of each eye (posteriorly); forebody wide in comparison to elytra; interval 3 of elytra without setiferous punctures; mentum tooth (Fig. 18) entire, acute apically. [Body length 12.3–20.2 mm; northern North Island] .. Genus Kupeplatynus Larochelle and Larivière, new genus 6 · April 30, 2021 Larochelle and Larivière 7(5). Mentum (Fig. 21) with numerous short setae (10–18) in addition to two long setae; circular foveae (Fig. 19) large. [Body length 10.0–12.3 mm; southern South Island] ............................... ...................................Genus Tuiplatynus Larochelle and Larivière, new genus — Mentum (Fig. 22) with two long setae only; circular foveae (Fig. 20) small. [Body length 7.0–13.9 mm; Three Kings Islands (TH), North Island, South Island, and Stewart Island] ..................... .........................................................Genus Ctenognathus Fairmaire Genus Platynus Bonelli, 1810 Fig. 37, 80, 159 Platynus Bonelli 1810: Tabula Synoptica. Type species. Carabus angusticollis Fabricius, 1801 (= Carabus assimilis Paykull, 1790), by subsequent monotypy in Germar (1817: 303). Description (New Zealand). Body length 13.0–14.2 mm. Color mostly dark. Metallic luster absent. Dorsal surface mostly glabrous. Forebody narrow in comparison to elytra. Head. Narrow. Mandibles moderately long. Labrum emarginate anteriorly. Eyes strongly convex, touching buccal fissures; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction deep dorsolaterally. Mentum tooth entire, rounded apically; two small circular foveae; two long setae. Submentum with four long setae. Palpi with terminal segment truncate apically. Ligula with two setae. Paraglossae glabrous. Thorax. Pronotum subquadrate, strongly cordate; two setiferous punctures (anterior and posterior) on each side. Prosternum glabrous apically, not compressed into a vertical ridge. Legs. Very long. Metacoxae with two posteroventral setae. Metafemora with two posteroventral setae. Protibiae with dorsal longitudinal groove; cleaning organ with two clip setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 strongly bilobed and asymmetrical apically. Metatarsomeres 5 setose ventrally. Elytra. Oblong. Free along suture (hindwings fully developed). Shoulders broad, rounded. Basal margin complete, reaching about scutellum. Sides widest about middle. Scutellar setiferous pore visible. Scutellar striole very long. Striae complete, consisting of impressed lines. Interval 3 with three setiferous punctures. Umbilicate series with 15–19 setiferous punctures. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view: moderately arcuate; basal lobe of basal bulb present; internal sac without scale-like sclerites. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to the left); basal bulb close to membranous area. References. Larochelle and Larivière 2001: 138–139 (catalogue), 2007: 82 (description, ecology, geographic distribution, references), 2016: 38 (list). Remark. This worldwide genus is represented by a single species, which occurs on the North Island, South Island, and Stewart Island. Platynus macropterus(Chaudoir, 1879) Fig. 37, 80, 159 Colpodes macropterus Chaudoir, 1879: 370. Holotype: male (MNHN) labeled “Ex Musaeo Chaudoir (typed) / HOLOTYPE (red label; hand-written) / macropterus Chaud. Nouv. Zélande Guerin (hand-written) / Holotype (red label; typed) / Holotype (typed) Colpodes macropterus Chd G.G. Perrault (hand-written) / G.G. Perrault det. 1988 (typed).” Anchomenus haasti Broun, 1882: 217 (redescribed in 1883: 217 and 1886: 747). Synonymized by Larochelle and Larivière (2001: 138). Ctenognathus macropterus: Hutton 1904: 144. Agonum (Platynus) haasti: Csiki 1931: 853. Colpodes haasti: Johns 1980: 62. Platynus macropterus: Liebherr 1998: 987, 997. Description. Body length 13.0–14.2 mm. Head, pronotum, and elytra piceous black; abdomen dark brown; antennal segments 1–4 piceous black, 5–11 reddish; palpi reddish; femora and tibiae piceous black; tarsi reddish. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 7 Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Iridescence absent. Very shiny. Head. Mandibles moderately curved anteriorly. Labrum moderately transverse, slightly emarginate anteriorly. Thorax. Pronotum subdepressed, impunctate, deeply wrinkled on disc and across base, narrow, widest before middle; apex slightly emarginate; anterolateral angles poorly developed, rounded; anterior bead complete; sides strongly rounded anteriorly, strongly sinuate posteriorly; lateral beads complete; lateral depressions moderately wide anteriorly, much widened posteriorly; posterolateral angles subrectangular; laterobasal foveae very deep and wide, prolonged forward; posterior bead incomplete, obsolete medially; base emarginate. Legs. Metatarsomeres 5 setose ventrally (with short setae). Elytra. Subdepressed, not sloping down toward apex, oblong. Sides slightly rounded. Scutellar striole impunctate. Striae deep, impunctate. Intervals subdepressed; interval 3 with three shallow, minute setiferous punctures. Subapical sinuations feeble. Aedeagus. Lateral view (Fig. 80): base strongly convex dorsally, with basal lobe narrow; middle moderately wide in apical half, moderately convex dorsally, strongly concave ventrally, with dorsal membranous area moderately wide in apical half and very long; apex strongly rounded dorsally, slightly concave ventrally, with extreme tip moderately wide and very long. Material examined. 157 specimens (AMNZ, BMNH, CMNH, JNNZ, LUNZ, MNHN, MONZ, NZAC). Geographic distribution (Fig. 159). North Island: AK, BP, CL, GB, HB, ND, RI, TK, TO, WN. South Island: BR, CO, DN, FD, KA, MB, MC, MK, NC, NN, OL, SD, SL, WD. Stewart Island. Ecology. Lowland, montane. Epigean, arboreal. Mostly wet open forests (broadleaf, podocarp, beech), swamp forests, shrublands, scrublands, forest edges, and tree plantations (pine); tussock grasslands and pastures. Associated with streams. Mostly open ground. Nocturnal; hides during the day under logs and stones. Biology. Seasonality: September–March, May, July–August. Tenerals: November–January, March. Predacious (based on mouthpart morphology). Predators: Feral cats. Occasionally infested with mites and fungi (Laboulbeniales). Defense mechanism: when alarmed, the adult emits a burnt chocolate-like smell. Dispersal power. Macropterous. Occasional flier to artificial lights. Swift runner. Occasional climber (on trees). Favored by human activities. Collecting techniques. Turning logs and stones. References. Larochelle and Larivière 2001: 138–139 (catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 114 (list), 2016: 38 (list). Genus Notagonum Darlington, 1952 Fig. 38–41, 81–84, 155–158 Notagonum Darlington, 1952: 127. Type species. Notagonum externum Darlington, 1952, by original designation. Description (New Zealand). Body length 7.3–10.4 mm. Color mostly dark. Metallic luster present. Dorsal surface mostly glabrous. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long. Labrum emarginate or subtruncate anteriorly. Eyes strongly convex, touching buccal fissures; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum tooth entire, acute apically; two small circular foveae; two long setae. Submentum with four long setae. Palpi with terminal segment obtuse apically. Ligula with two setae. Paraglossae glabrous. Thorax. Pronotum transverse, cordate, subquadrate, or rectangular; two setiferous punctures (anterior and posterior) on each side. Prosternum glabrous apically, not compressed into a vertical ridge. Legs. Moderately long (very long in N. submetallicum). Metacoxae with two posteroventral setae. Metafemora with two posteroventral setae. Protibiae with dorsal longitudinal groove; cleaning organ with two clip setae. Mesoand metatarsomeres 1–4 poorly developed, not carinate dorsally, shallowly bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous or setose ventrally. Elytra. Oblong (pear-shaped in N. chathamense). Free along suture (hindwings fully developed) or fused (hindwings about half developed in N. chathamense). Basal margin complete, reaching about scutellum. Shoulders broad (narrow in N. chathamense), rounded. Sides widest behind middle. Scutellar setiferous pore visible. Scutellar striole very long. Striae complete, consisting of 14 · April 30, 2021 Larochelle and Larivière developed, rounded; anterior bead incomplete, obsolete medially; sides slightly rounded anteriorly, not sinuate posteriorly; lateral beads incomplete, obsolete around posterolateral angles; lateral depressions narrow anteriorly, widened posteriorly; posterolateral angles obtusely rounded; laterobasal foveae very deep and wide, rounded, not prolonged forward; posterior bead absent; base emarginate. Legs. Moderately long. Protibiae without dorsal longitudinal groove. Elytra. Slightly convex, not sloping down toward apex (sloping down in other species), oblong (subovate in other species), widest about middle. Basal margin incomplete, reaching about stria 3. Shoulders moderately rounded. Sides slightly rounded. Scutellar striole very long, impunctate. Striae shallow, finely punctate. Intervals moderately to strongly convex; interval 3 with two obsolete setiferous punctures. Umbilicate series with 23–24 setiferous punctures. Subapical sinuations moderately strong. Apices truncate-emarginate. Abdomen. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 85): moderately arcuate, stout; base strongly convex dorsally, with basal lobe moderately wide; middle subparallel, slightly convex dorsally, slightly concave ventrally, with dorsal membranous area short, moderately wide toward base dorsally; apex moderately concave dorsally, strongly concave ventrally, with extreme tip narrow, very long, and strongly curved downward. Dorsal view: moderately wide, symmetrical (ostium of membranous area dorsal). Material examined. 26 specimens (AMNZ, CMNZ, LUNZ, NZAC). Geographic distribution (Fig. 160). North Island: AK, CL, ND. Ecology. Lowland. Fossorial, amphibious. Margins of rills (about 1–2 m wide) crossing dense wet forests (broadleaf, podocarp). Associated with gullies and ravines. Shaded ground; wet, stony, gravelly or muddy soil. Nocturnal; hides during the day under embedded stones. Biology. Seasonality: December–July. Tenerals: February–March, June–July. Predacious (based on mouthpart morphology). Defense mechanism: when alarmed, the adult escapes by running and diving into the water. Dispersal power. Subapterous (incapable of flight). Moderate runner. Excellent swimmer. Collecting techniques. Turning embedded stones; pitfall trapping. Remarks. The species is named after a geographic area (Mangamuka Gorge) where the beetle is especially abundant, and is applied as a noun in apposition. In addition to diagnostic characters of the male genitalia, Prosphodrus mangamuka has the following distinguishing features: large and robust, body length 23.0–26.2 mm; antennae moderately long; eyes moderately large and convex; terminal segment of palpi obtuse; pronotum subquadrate; legs moderately long; elytra oblong, not sloping down toward apex, apices truncate-emarginate. Prosphodrus mangamuka occurs in northern areas of the North Island (ND, AK, CL). Prosphodrus waltoni Britton, 1959 Fig. 43, 86, 164 Prosphodrus waltoni Britton, 1959: 106. Holotype: male (NZAC) labeled “Type (circular red-bordered label; typed) / alive on wall 500 yrds inside Waipuna Cave Te Kuiti [WO] 11.Jan.1958. B.M. May (hand-written) / HOLOTYPE [male symbol] Prosphodrus waltoni gen. sp. n. (hand-written).” Paratype: one female (BMNH) from the same locality as the holotype, bearing a blue paratype label. Description. Body length 17.0–20.1 mm. Head, pronotum, and elytra black; abdomen black or reddish brown; antennae and palpi reddish brown; legs dark brown. Microsculpture isodiametric on head, moderately transverse on pronotum and elytra. Very shiny, without metallic luster. Head. Mandibles much longer than in other species, strongly curved anteriorly. Labrum strongly transverse, slightly biconcave anteriorly. Antennae very long. Eyes small, strongly convex. Tempora strongly inflated and very long (about two times as long as eyes). Mentum with six setae. Submentum with eight setae. Palpi with terminal segment truncate apically. Thorax. Pronotum strongly convex, impunctate, feebly wrinkled medially and laterally, wide, strongly cordate, widest before middle; apex slightly emarginate; anterolateral angles moderately developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded anteriorly, strongly sinuate posteriorly; lateral beads incomplete, obsolete in basal third; lateral depressions complete, widened anteriorly and posteriorly; posterolateral angles strongly rounded; laterobasal foveae very deep and wide, elongate (reaching pronotal apex), prolonged forward; posterior Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 15 bead absent; base emarginate. Legs. Very long. Protibiae with dorsal longitudinal groove. Elytra. Moderately convex, sloping down toward apex, subovate, widest about middle. Basal margin complete, reaching about scutellum. Shoulders strongly rounded. Sides moderately rounded. Scutellar striole very long, impunctate. Striae deep, finely punctate. Intervals slightly convex; interval 3 with three obsolete setiferous punctures. Umbilicate series with 19–20 setiferous punctures. Subapical sinuations moderately strong. Apices obtusely rounded. Abdomen. Sternum VII (last visible sternum): both sexes with two long apical ambulatory setae. Aedeagus. Lateral view (Fig. 86): strongly arcuate in basal half; base slightly convex dorsally, with basal lobe moderately wide; middle very wide, strongly convex, slightly concave toward apex dorsally, slightly convex ventrally, with dorsal membranous area strongly widened in apical half; apex subtriangular, moderately concave dorsally, almost straight, slightly convex ventrally, with extreme tip narrow and rather short. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left). Material examined. 37 specimens (AMNZ, BMNH, CMNH, CMNZ, JNNZ, MONZ, NZAC). Geographic distribution (Fig. 164). North Island: AK, CL, RI, TK, WO. Ecology. Lowland, montane. Fossorial, amphibious. Margins of rills (about 1–2 m wide) and seepages crossing dense wet forests (broadleaf, podocarp); caves (occasionally). Associated with gullies and ravines. Shaded ground; wet, stony or gravelly soil. Nocturnal; hides during the day under embedded stones, logs, and clods of clay. Gregarious. Biology. Seasonality: January–February, April, June–July. Tenerals: March–April, June. Predacious (based on mouthpart morphology). Defense mechanism: when alarmed, the adult escapes by running and diving into the water. Dispersal power. Subapterous (incapable of flight). Swift runner. Excellent swimmer. Collecting techniques. Turning embedded stones and logs, also clods of clay; pitfall trapping. References. Larochelle and Larivière 2001: 139 (catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 114 (list), 2016: 38 (list). Prosphodrus sirvidi Larochelle and Larivière, new species Fig. 44, 87, 162 Prosphodrus sirvidi Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “[NEW ZEALAND WN] Wellington (typed) Akatarawa Range 1300’ 11.5.73 (hand-written) J.I. Townsend (typed) / J.I. Townsend Collection (typed) / HOLOTYPE [male symbol] Prosphodrus sirvidi Larochelle & Larivière, 2021 (red label; typed).” Paratypes: two females (CMNZ, NZAC) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 14.0–16.1 mm. Head, pronotum, and elytra dark brown; abdomen dark rufous; antennae, palpi, and legs reddish. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Moderately shiny, with metallic luster (bronze). Head. Mandibles strongly curved anteriorly. Labrum moderately transverse, slightly to moderately emarginate anteriorly. Antennae very long. Eyes small, strongly convex. Tempora moderately inflated and long (about half as long as eyes). Mentum with two setae. Submentum with six setae. Palpi with terminal segment truncate apically. Thorax. Pronotum feebly convex, impunctate, deeply wrinkled throughout, narrow (wide in other species), subcordate, widest before middle; apex subtruncate; anterolateral angles well developed, subangulate; anterior bead incomplete, present only around anterolateral angles; sides angulate anteriorly, moderately sinuate posteriorly; lateral beads complete, widened basally; lateral depressions absent anteriorly, wide posteriorly; posterolateral angles rectangular; laterobasal foveae moderately deep and wide, elongate (reaching anterolateral setiferous punctures), prolonged forward; posterior bead obsolete; base emarginate. Legs. Very long. Protibiae with dorsal longitudinal groove obsolete. Elytra. Subdepressed, sloping down toward apex, subovate, widest behind middle. Basal margin incomplete, reaching about stria 3. Shoulders angulate. Sides slightly rounded. Scutellar striole short or moderately long, impunctate. Striae deep, finely punctate. Intervals depressed; interval 3 with or without a single obsolete setiferous puncture. Umbilicate series with 17–18 setiferous punctures. Subapical sinuations feeble. Apices broadly rounded. Abdomen. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory 16 · April 30, 2021 Larochelle and Larivière setae. Aedeagus. Lateral view (Fig. 87): strongly arcuate, transversely wrinkled in apical third; base slightly convex dorsally, with basal lobe very wide; middle strongly convex, slightly concave toward apex dorsally, strongly concave ventrally, without visible dorsal membranous area; apex subtriangular, slightly concave dorsally and ventrally, with extreme tip narrow and very long. Dorsal view: narrow, symmetrical (ostium of membranous area dorsal). Material examined. 42 specimens (CMNZ, JNNZ, MONZ, NZAC). Geographic distribution (Fig. 162). North Island: HB, RI, WA, WI, WN. Ecology. Lowland, montane. Fossorial, amphibious. Margins of rills (about 1 m wide) and seepages crossing dense wet forests (beech, broadleaf, podocarp); caves (occasionally). Associated with gullies and ravines. Shaded ground; wet clay soil. Nocturnal; hides during the day under embedded stones. Gregarious. Biology. Seasonality: September, November, February–May, August. Predacious (based on mouthpart morphology). Defense mechanism: when alarmed, the adult escapes by running and diving into the water. Dispersal power. Subapterous (incapable of flight). Swift runner. Excellent swimmer. Collecting technique. Turning embedded stones. Remarks. The species is named after our friend and colleague Phil Sirvid (Museum of New Zealand Te Papa Tongarewa, Wellington) for his special help and encouragement in our entomological studies. In addition to diagnostic characters of the male genitalia, Prosphodrus sirvidi has the following distinguishing features: antennae, palpi, and legs reddish; pronotum subcordate and narrow, sides moderately sinuate posteriorly, anterolateral angles subangulate. Prosphodrus sirvidi occurs in southern areas of the North Island (RI–HB to WN–WA), mostly in the Wellington and Wairarapa regions. Prosphodrus waimana Larochelle and Larivière, new species Fig. 45, 88, 163 Prosphodrus waimana Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “NEW ZEALAND BP Urewera NP, Waimana Vly, Onepu Hut tk [=track] 1.XII.1995 Larivière, Larochelle (typed) / Shaded edges, swift rill; stony-rocky; broadleaf for. [=forest] Under ½ emerged stones (typed) / HOLOTYPE [male symbol] Prosphodrus waimana Larochelle & Larivière, 2021 (red label; typed).” Paratypes: one male (CMNZ) from the same locality as the holotype and one female (NZAC) from Urewera National Park, Waimana Valley, Te Waiiti Stream (BP), bearing blue paratype labels. Description. Body length 16.1–18.0 mm. Head, pronotum, and elytra black; abdomen dark brown; antennae and palpi reddish brown; legs dark brown. Microsculpture moderately transverse on head, pronotum, and elytra. Very shiny, without metallic luster. Head. Mandibles slightly curved anteriorly. Labrum moderately transverse, slightly to moderately emarginate anteriorly. Antennae very long. Eyes small, strongly convex. Tempora moderately inflated and long (about half as long as eyes). Mentum with two setae. Submentum with four setae. Palpi with terminal segment truncate apically. Thorax. Pronotum moderately convex, impunctate, feebly wrinkled medially and laterally, wide, trapezoid, widest before middle; apex subtruncate; anterolateral angles moderately developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded anteriorly, not sinuate posteriorly; lateral beads complete, widened basally; lateral depressions narrow anteriorly, widened posteriorly; posterolateral angles subrectangular; laterobasal foveae very deep and wide, rounded, not prolonged forward; posterior bead complete; base emarginate. Legs. Very long. Protibiae with dorsal longitudinal groove obsolete. Elytra. Moderately convex, sloping down toward apex, subovate, widest behind middle. Basal margin incomplete, reaching about stria 4. Shoulders angulate. Sides slightly rounded. Scutellar striole very long, impunctate. Striae deep (shallower at base), finely punctate. Intervals depressed; interval 3 with or without one or two obsolete setiferous punctures. Umbilicate series with 19–20 setiferous punctures. Subapical sinuations absent. Apices obtuse. Abdomen. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 88): strongly arcuate in basal half; base slightly convex dorsally, with basal lobe moderately wide; middle subparallel, strongly convex in basal half, almost straight in apical half dorsally, moderately concave ventrally, without visible dorsal membranous area; Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 17 apex subtriangular, strongly concave dorsally, slightly concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, symmetrical (ostium of membranous area dorsal). Material examined. 147 specimens (CMNZ, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 163). North Island: BP, GB, TO. Ecology. Lowland, montane. Fossorial, amphibious. Margins of rills (about 1–2 m wide) and seepages crossing dense wet forests (broadleaf, podocarp). Associated with gullies and ravines. Shaded ground; wet, stony or gravelly soil. Nocturnal; hides during the day under embedded stones. Gregarious. Biology. Seasonality: September–April, July. Tenerals: November, January–February. Predacious (based on mouthpart morphology). Defense mechanism: when alarmed, the adult escapes by running and diving into the water. Dispersal power. Subapterous (incapable of flight). Swift runner. Excellent swimmer. Collecting techniques. Turning embedded stones; pitfall trapping. Remarks. The species is named after the type locality Waimana Valley (BP), and is applied as a noun in apposition. Prosphodrus waimana is morphologically close to P. occultus. In addition to diagnostic characters of the male genitalia, P. waimana has the following distinguishing features: pronotum feebly wrinkled medially and laterally, posterior bead complete. Both species occur in eastern areas of the North Island although they are allopatric: P. waimana is present in BP, TO, and GB (East Cape area), while P. occultus is found in RI, HB, and GB (excluding the East Cape area). Prosphodrus occultus Britton, 1960 Fig. 46, 89, 161 Prosphodrus occultus Britton, 1960: 123. Holotype: male (NZAC) labeled “Type (circular red-bordered label; typed) / in debris 400 yrds from the entrance in Simpsons Cave Wairoa [GB] Hawkes Bay Sept. 1959 A. Norgrove (hand-written) / HOLOTYPE [male symbol] Prosphodrus occultus mihi E.B. Britton det. 1959 (hand-written).” Paratype: one female (BMNH) from the same locality as the holotype, bearing a blue paratype label. Description. Body length 16.3–19.0 mm. Head, pronotum, elytra black; abdomen dark brown; antennae and palpi reddish brown; legs dark brown. Microsculpture moderately transverse on head, pronotum, and elytra. Very shiny, without metallic luster. Head. Mandibles slightly curved anteriorly. Labrum moderately transverse, slightly to moderately emarginate anteriorly. Antennae very long. Eyes small, strongly convex. Tempora moderately inflated and long (about half as long as eyes). Mentum with two setae. Submentum with four setae. Palpi with terminal segment truncate apically. Thorax. Pronotum moderately convex, impunctate, deeply wrinkled throughout, wide, trapezoid, widest before middle; apex subtruncate or slightly emarginate; anterolateral angles moderately developed, rounded; anterior bead incomplete, obsolete medially; sides moderately rounded anteriorly, not sinuate posteriorly; lateral beads complete, widened basally; lateral depressions narrow anteriorly, widened posteriorly; posterolateral angles subrectangular; laterobasal foveae very deep and wide, rounded, not prolonged forward; posterior bead obsolete medially; base emarginate. Legs. Very long. Protibiae with dorsal longitudinal groove obsolete. Elytra. Moderately convex, sloping down toward apex, subovate, widest behind middle. Basal margin incomplete, reaching about stria 4. Shoulders angulate. Sides slightly rounded. Scutellar striole moderately long or very long, impunctate. Striae deep (although shallower at base), finely punctate. Intervals depressed; interval 3 with or without one or two obsolete setiferous punctures. Umbilicate series with 19–21 setiferous punctures. Subapical sinuations absent. Apices obtuse or obtusely rounded. Abdomen. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 89): strongly arcuate medially, transversely wrinkled in apical third; base slightly convex dorsally, with basal lobe very wide; middle strongly convex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and long; apex slender, sinuate dorsally, slightly concave ventrally, with extreme tip narrow and very long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left). Material examined. 85 specimens (AMNZ, BMNH, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 161). North Island: GB, HB, RI. 18 · April 30, 2021 Larochelle and Larivière Ecology. Lowland, montane. Fossorial, amphibious. Margins of rills (about 1–2 m wide) and seepages crossing dense wet forests (broadleaf, podocarp); caves (occasionally). Associated with gullies and ravines. Shaded ground; wet stony soil. Nocturnal; hides during the day under embedded stones. Gregarious. Biology. Seasonality: October–May. Tenerals: January–February. Predacious (based on mouthpart morphology). Defense mechanism: when alarmed, the adult escapes by running and diving into the water. Dispersal power. Subapterous (incapable of flight). Swift runner. Excellent swimmer. Collecting technique. Turning embedded stones. References. Larochelle and Larivière 2001: 139 (catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 114 (list), 2016: 38 (list). Genus Maoriplatynus Larochelle and Larivière, new genus Fig. 47, 90, 154 Type species. Maoriplatynus marrisi new species, by present designation. Description. Body length 12.5–12.7 mm. Color mostly dark. Metallic luster absent. Dorsal surface mostly glabrous. Forebody narrow in comparison to elytra. Head. Very wide. Mandibles very long. Labrum subtruncate anteriorly. Eyes moderately convex, separated from buccal fissures; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction deep dorsolaterally. Mentum tooth bifid apically (as in Prosphodrus); two large circular foveae; two long setae. Submentum with twelve short setae in addition to two long setae. Palpi with terminal segment obtuse apically. Ligula with two setae. Paraglossae glabrous. Thorax. Pronotum cordate; a single setiferous puncture on each side (anteriorly). Prosternum glabrous apically, not compressed into a vertical ridge. Legs. Very long. Metacoxae with two posteroventral setae. Metafemora with two or three posteroventral setae. Protibiae with dorsal longitudinal groove; cleaning organ with two clip setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 strongly bilobed and asymmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Ovate. Fused along suture (hindwings vestigial). Basal margin complete, reaching about scutellum. Shoulders narrow, rounded. Sides widest about middle. Scutellar setiferous pore visible. Scutellar striole short. Striae complete, consisting of impressed lines. Interval 3 with three setiferous punctures. Umbilicate series with 17–20 setiferous punctures. Apices mucronate, turned upward (not turned upward in other genera). Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sterna V–VI: both sexes with numerous short setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae and numerous short setae; female with four long apical ambulatory setae and numerous short setae. Aedeagus. Lateral view: slightly arcuate; basal lobe of basal bulb absent; internal sac without scale-like sclerites. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Remarks. The generic name is derived from Maori (the Polynesian people who colonized New Zealand) and Platynus (the type genus of the tribe Platynini), genus masculine. This monotypic genus appears to be a genetically highly distinctive taxon characterized as follows: abdominal sterna V–VII of both sexes with numerous short setae in addition to two long apical ambulatory setae; submentum with twelve short setae in addition to two long setae; elytral apices turned upward; aedeagus (in lateral view) slightly arcuate. This genus is restricted to the Three Kings Islands (TH). Maoriplatynus marrisi Larochelle and Larivière, new species Fig. 47, 90, 154 Maoriplatynus marrisi Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “[NEW ZEALAND TH] Three Kings Is Great I. Nov. 70 NZ. Ent. Div. Exp. (typed) / HOLOTYPE [male symbol] Maoriplatynus marrisi Larochelle & Larivière, 2021 (red label; typed).” Paratypes: one female (NZAC) and one male (LUNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 12.5–12.7 mm. Head, pronotum, and elytra black; abdomen dark brown; antennae, palpi, and tarsi reddish; femora and tibiae rufopiceous. Microsculpture absent on head, very transverse (with Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 19 microlines) on pronotum and elytra. Iridescence present on pronotum and elytra. Very shiny. Head. Mandibles slightly curved anteriorly. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc and across base, narrow, strongly cordate, widest before middle; apex strongly emarginate; anterolateral angles well developed, angulate; anterior bead incomplete, obsolete medially; sides strongly rounded, strongly sinuate posteriorly; lateral beads incomplete, obsolete anteriorly; lateral depressions narrow; posterolateral angles rectangular, acute at apex; laterobasal foveae shallow, moderately wide, prolonged forward; posterior bead obsolete; base emarginate. Elytra. Slightly convex, not sloping down toward apex. Shoulders broadly rounded. Sides strongly rounded. Scutellar striole impunctate. Striae deep, impunctate. Intervals moderately convex; interval 3 with three moderately deep, small setiferous punctures. Subapical sinuations feeble. Aedeagus. Lateral view (Fig. 90): slender, slightly arcuate; base slightly convex, almost straight dorsally; middle slightly convex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and very long; apex slender, moderately convex dorsally, slightly concave ventrally, with extreme tip moderately wide and long. Material examined. 30 specimens (CMNZ, LUNZ, NZAC). Geographic distribution (Fig. 154). Offshore Islands: TH–Great Island, Tasman Valley. Ecology. Coastal lowland. Epigean. Wet forests (broadleaf). Associated with streams. Shaded ground; wet soil. Nocturnal; active at night on mossy stream banks. Gregarious. Biology. Seasonality: October–December. Tenerals: December. Predacious (based on mouthpart morphology). Regularly infested with mites. Dispersal power. Subapterous (incapable of flight). Swift runner. Collecting technique. Hand-collecting. Remarks. This species is named after our friend and colleague John M.W. Marris (Lincoln University, Lincoln, Christchurch) for his special help and encouragement in our entomological studies. In addition to generic characters, Maoriplatynus marrisi has the following distinguishing features: iridescence present on pronotum and elytra; pronotum narrow, strongly cordate; elytral interval 3 with three setiferous punctures. Genus Kiwiplatynus Larochelle and Larivière, new genus Fig. 48–49, 91–92, 149–150 Type species. Colpodes bidens Chaudoir, 1878, by present designation. Description. Body length 16.0–19.3 mm. Color mostly dark. Metallic luster absent. Dorsal surface mostly glabrous. Forebody narrow in comparison to elytra. Head. Narrow. Mandibles moderately long. Labrum truncate or subtruncate anteriorly. Eyes strongly convex, separated from buccal fissures; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction deep dorsolaterally. Mentum tooth emarginate apically; two small circular foveae; two long setae. Submentum with six or eight long setae. Palpi with terminal segment truncate apically. Ligula with two setae. Paraglossae glabrous. Thorax. Pronotum cordate; a single setiferous puncture on each side (anteriorly). Prosternum glabrous apically, not compressed into a vertical ridge. Legs. Very long. Metacoxae with three or four posteroventral setae. Metafemora with three posteroventral setae. Protibiae with dorsal longitudinal groove; cleaning organ with three clip setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and symmetrical apically. Metatarsomeres 5 setose ventrally. Elytra. Subovate. Fused along suture (hindwings vestigial). Basal margin complete, reaching about scutellum. Shoulders narrow, rounded. Sides widest about middle. Scutellar setiferous pore visible. Scutellar striole short or moderately long. Striae complete, consisting of impressed lines. Interval 3 with a single subapical setiferous puncture or two setiferous punctures (anterior puncture absent). Umbilicate series with 22–25 setiferous punctures. Apices mucronate or obtusely triangular. Abdomen. Sternum IV: both sexes with two to four long ambulatory setae. Sterna V–VI: both sexes with three to five long apical ambulatory setae. Sternum VII (last visible sternum): male with two or four long apical ambulatory setae; female with six or eight long apical ambulatory setae. Aedeagus. Lateral view: moderately arcuate; basal lobe of basal bulb present; internal sac with scale-like sclerites. Dorsal view: moderately wide or very wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb close to membranous area. 20 · April 30, 2021 Larochelle and Larivière Remarks. The generic name is derived from Kiwi (the vernacular name of an ancient group of New Zealand birds; also a major national symbol) and Platynus (the type genus of the tribe Platynini), genus masculine. The morphological characters unifying species of Kiwiplatynus are: forebody narrow in comparison to elytra; two setiferous punctures on inner side of each eye; mentum tooth emarginate apically; metatarsomeres 4 symmetrical apically; interval 3 of elytra with one or two setiferous punctures; abdominal sterna V–VI of both sexes with three to five long apical ambulatory setae; aedeagus (in lateral view) moderately arcuate; internal sac of aedeagus with scalelike sclerites. Furthermore, three morphological characters are shared between Kiwiplatynus and Kupeplatynus that set them apart from Ctenognathus and Tuiplatynus: cleaning organ of protibiae with three clip setae; neck constriction deep dorsolaterally; basal bulb of aedeagus close to membranous area. Kiwiplatynus species occur mostly in northern areas of the North Island. Key to species of Kiwiplatynus 1. Elytra with striae impunctate, interval 3 with two setiferous punctures (medially and subapically); submentum with six setae; sternum VII (last visible abdominal sternum), male with two long apical ambulatory setae, female with six long apical ambulatory setae; Fig. 48. [Body length 16.0–19.3 mm; North Island (ND to TK–GB; WN)] ........................ Kiwiplatynus bidens (Chaudoir) — Elytra with striae coarsely punctate, interval 3 with a single setiferous puncture (subapically); submentum with eight setae; sternum VII, male with four long apical ambulatory setae, female with eight long apical ambulatory setae; Fig. 49. [Body length 16.1–19.0 mm; central North Island (WO, TK, TO)] ......................... Kiwiplatynus taranaki Larochelle and Larivière, new species Kiwiplatynus bidens (Chaudoir, 1878), new combination Fig. 48, 91, 149 Colpodes bidens Chaudoir, 1878: 303. Holotype: female (MNHN) labeled “ExMusaeo Chaudoir (typed) / bidens Chaud. Nouv. Zélande (hand-written).” Anchomenus batesi Broun, 1880: 21. Synonymized by Watt (1961: 93). Ctenognathus bidens: Broun 1893: 986. Ctenognathus batesi: Broun 1893: 986. Description. Body length 16.0–19.3 mm. Head, pronotum, elytra, and abdomen black; antennal segments 1–3 rufopiceous, segments 4–11 reddish; palpi rufopiceous; legs dark rufous. Microsculpture isodiametric on head, very transverse (with microlines) on pronotum, and isodiametric on elytra. Iridescence present on pronotum. Very shiny on head and pronotum, moderately shiny on elytra. Head. Mandibles moderately curved anteriorly. Labrum moderately transverse, truncate anteriorly. Mentum: medial tooth moderately emarginate apically. Submentum with six setae. Thorax. Pronotum slightly convex, obsoletely punctate across base, obsoletely wrinkled on disc and across base, narrow, slightly cordate, widest before middle; apex slightly emarginate; anterolateral angles well developed, angulately rounded; anterior bead complete; sides moderately rounded, not sinuate posteriorly; lateral beads complete; lateral depressions wide, more so posteriorly; posterolateral angles obtuse; laterobasal foveae very deep and wide, prolonged forward; posterior bead complete, less impressed near posterolateral angles; base subtruncate. Legs. Metatarsomeres 5 setose ventrally (with long setae). Elytra. Slightly convex, not sloping down toward apex. Shoulders moderately rounded. Sides moderately rounded. Scutellar striole moderately long, impunctate. Striae deep, impunctate. Intervals moderately convex; interval 3 with two shallow, minute setiferous punctures (medially and subapically). Umbilicate series with 24–25 setiferous punctures. Subapical sinuations moderately strong. Apices mucronate internally, rounded externally. Abdomen. Sternum IV: both sexes with two or three long apical ambulatory setae. Sternum V: both sexes with three or four long apical ambulatory setae. Sternum VI: both sexes with four long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with six long apical ambulatory setae. Aedeagus. Lateral view (Fig. 91): moderately arcuate; base moderately convex dorsally, with basal lobe rather narrow; middle subparallel, straight dorsally, moderately concave ventrally, with dorsal membranous area strongly widened from base to apex and very long; apex subtriangular, slightly convex dorsally, rather straight ventrally, with extreme tip narrow, short, Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 21 and moderately curved downward; internal sac with a few scale-like sclerites. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb close to membranous area. Material examined. 2245 specimens (AMNZ, BMNH, CMNH, JNNZ, LUNZ, MNHN, MONZ, NZAC). Geographic distribution (Fig. 149). North Island: AK, BP, CL, GB, ND, TK, TO, WN, WO. Ecology. Lowland, montane, subalpine. Epigean, arboreal. Mesic forests (podocarp, broadleaf), tree plantations (pine), and shrublands. Shaded ground; moderately moist soil. Nocturnal; hides during the day under logs, fallen trees and branches, or under stones. Gregarious. Biology. Seasonality: Throughout the year. Mating observed in October (BP). Tenerals: December–March. Predacious (based on mouthpart morphology). Predators: Kiwi birds. Regularly infested with fungi (Laboulbeniales). Occasionally infested with mites. Dispersal power. Subapterous (incapable of flight). Swift runner. Frequent climber (on trees). Collecting techniques. Pitfall trapping; turning logs, fallen trees and branches, as well as stones. References. Larochelle and Larivière 2001: 133–134 (as Ctenognathus bidens; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 111 (as C. bidens; list), 2016: 36 (as C. bidens; list). Kiwiplatynus taranaki Larochelle and Larivière, new species Fig. 49, 92, 150 Kiwiplatynus taranaki Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “NEW ZEALAND TK Mt Messenger For [=Forest], Mokau Rd-Kiwi Rd tk [=track] 200m 385350S 1743600E 5–11.XII.1999 Larivière, Larochelle (typed) / Wet broadleaf forest: pittraps. (typed) / HOLOTYPE [male symbol] Kiwiplatynus taranaki Larochelle & Larivière, 2021 (red label; typed).” Paratypes: one female (NZAC) and one male (AMNZ) from Mount Messenger Forest, Mokau Road-White Cliffs track, bearing blue paratype labels. Description. Body length 16.1–19.0 mm. Head, pronotum, and elytra black; abdomen blackish; antennae, palpi, and legs rufopiceous. Microsculpture very transverse (with microlines) on head and pronotum, moderately transverse on elytra. Iridescence present on head and pronotum. Very shiny on head and pronotum, moderately shiny on elytra. Head. Mandibles moderately curved anteriorly. Labrum moderately transverse, subtruncate anteriorly. Mentum: medial tooth strongly emarginate apically. Submentum with eight setae. Thorax. Pronotum slightly convex, obsoletely punctate, obsoletely wrinkled on disc, narrow, moderately cordate, widest before middle; apex slightly emarginate; anterolateral angles well developed, broadly rounded; anterior bead complete; sides moderately rounded and sinuate posteriorly; lateral beads complete; lateral depressions narrow; posterolateral angles rectangular, acute at tip; laterobasal foveae moderately deep, very wide, prolonged forward; posterior bead absent; base subtruncate. Legs. Metatarsomeres 5 setose ventrally (with long setae). Elytra. Slightly convex, not sloping down toward apex. Shoulders moderately rounded. Sides moderately rounded. Scutellar setiferous pore visible. Scutellar striole short or moderately long, punctate. Striae deep, coarsely punctate. Intervals strongly convex; interval 3 with a single shallow, minute subapical setiferous puncture. Umbilicate series with 22–23 setiferous punctures. Subapical sinuations feeble. Apices obtusely triangular. Abdomen. Sternum IV: both sexes with four long apical ambulatory setae. Sterna V–VI: both sexes with four or five long apical ambulatory setae. Sternum VII (last visible sternum): male with four long apical ambulatory setae; female with eight long apical ambulatory setae. Aedeagus. Lateral view (Fig. 92): moderately arcuate; base strongly convex dorsally, with basal lobe rather narrow; middle subparallel, slightly biconcave dorsally, slightly concave ventrally, with dorsal membranous area moderately widened from base to apex and moderately long; apex subtriangular, moderately biconvex dorsally, slightly concave ventrally, with extreme tip narrow, moderately long, and moderately curved downward; internal sac with numerous scale-like sclerites. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb close to membranous area. Material examined. 65 specimens (AMNZ, BMNH, LUNZ, MONZ, NZAC). 22 · April 30, 2021 Larochelle and Larivière Geographic distribution (Fig. 150). North Island: TK, TO, WO. Ecology. Lowland, montane, subalpine. Epigean. Wet forests (broadleaf, beech, podocarp) and swamp forests. Associated with streams and slopes. Shaded ground; wet soil. Nocturnal; hides during the day under logs. Gregarious. Biology. Seasonality: October–January, March–April. Tenerals: November–December. Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Swift runner. Collecting techniques. Pitfall trapping; turning logs. Remarks. This species is named after the Taranaki region which includes the type locality Mount Messenger (TK), and is applied as a noun in apposition. Kiwiplatynus taranaki is morphologically close to K. bidens. In addition to diagnostic characters of the male genitalia, K. taranaki has the following distinguishing features: submentum with eight setae; elytra with striae coarsely punctate, interval 3 with a single subapical setiferous puncture; sternum VII, male with four long apical ambulatory setae, female with eight long apical ambulatory setae. Kiwiplatynus taranaki is known from the central North Island (WO, TK, TO), while K. bidens occurs throughout the North Island (ND to TK–GB; WN). Genus Kupeplatynus Larochelle and Larivière, new genus Fig. 50–52, 93–95, 151–153 Type species. Colpodes crenatus Chaudoir, 1878, by present designation. Description. Body length 12.8–20.2 mm. Color mostly dark. Metallic luster absent. Dorsal surface mostly glabrous. Forebody very wide in comparison to elytra. Head. Moderately wide or very wide. Mandibles moderately long or very long. Labrum subtruncate or emarginate anteriorly. Eyes moderately convex, separated from buccal fissures; a single setiferous puncture on inner side of each eye (posteriorly). Tempora inflated or not. Neck constriction deep dorsolaterally. Mentum tooth entire, acute apically; two small circular foveae; two long setae. Submentum with two, four, or six long setae. Palpi with terminal segment truncate apically. Ligula with two setae. Paraglossae glabrous. Thorax. Pronotum cordate or suborbicular; a single setiferous puncture on each side (anteriorly). Prosternum glabrous or pubescent apically, not compressed into a vertical ridge. Legs. Moderately long or very long. Metacoxae with three or four posteroventral setae. Metafemora with two to four posteroventral setae. Protibiae with dorsal longitudinal groove; cleaning organ with three clip setae. Mesoand metatarsomeres 1–4 poorly or well developed, tricarinate or not carinate dorsally, shallowly or deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, moderately or strongly asymmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Oblong, ovate, or subovate. Fused along suture (hindwings vestigial). Basal margin complete (reaching about scutellum) or incomplete (reaching about stria 1). Shoulders narrow, rounded or obtuse. Sides widest about middle. Scutellar setiferous pore visible or not. Scutellar striole short. Striae complete, consisting of impressed lines. Interval 3 without setiferous punctures. Umbilicate series with 17–25 setiferous punctures. Apices rounded or mucronate. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four or six long apical ambulatory setae. Aedeagus. Lateral view: strongly arcuate, semicircular; basal lobe of basal bulb present; internal sac with or without scale-like sclerites. Dorsal view: moderately wide, symmetrical (ostium of membranous area dorsal) or asymmetrical (ostium of membranous area deflected to the left); basal bulb close to membranous area. Remarks. The generic name is derived from Kupe (the legendary Polynesian navigator to whom is attributed the discovery of New Zealand) and Platynus (the type genus of the tribe Platynini), genus masculine. The morphological characters unifying species of Kupeplatynus are: forebody wide in comparison to elytra; a single setiferous puncture on inner side of each eye (posteriorly); mentum tooth entire, acute apically; metatarsomeres 4 asymmetrical apically; interval 3 of elytra without setiferous punctures; abdominal sterna IV–VI of both sexes with two long apical ambulatory setae; aedeagus (in lateral view) strongly arcuate, semicircular. Kupeplatynus species occur in northern areas of the North Island. See also Remarks under Kiwiplatynus. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 23 Key to species of Kupeplatynus 1. Femora and tibiae dark rufous; antennae reddish; elytra oblong, scutellar setiferous pore invisible, striae impunctate, intervals depressed; Fig. 50. [Body length 12.8–14.0 mm; submentum with six setae; apex of prosternum pubescent; northern North Island (ND to WO)] .......................... ........................................................Kupeplatynus sulcitarsis (Broun) — Femora and tibiae black piceous; antennal segments 1–3 rufopiceous, 4–11 pale reddish; elytra subovate or ovate, scutellar setiferous pore visible, striae coarsely punctate, intervals convex ............ 2 2(1). Mandibles very long; submentum with two setae; pronotum slightly convex, moderately cordate, widest before middle, sides moderately sinuate posteriorly; apex of prosternum pubescent; elytra subovate; Fig. 51. [Body length 16.5–20.2 mm; northern North Island (ND to TK–BP)] .................. ......................................................Kupeplatynus crenatus (Chaudoir) — Mandibles moderately long; submentum with four setae; pronotum strongly convex, suborbicular, widest about middle, sides not sinuate posteriorly; apex of prosternum glabrous; elytra ovate; Fig. 52. [Body length 15.3–18.4 mm; northern North Island (ND to TK–TO)] ......................... .........................................................Kupeplatynus lucifugus (Broun) Kupeplatynus sulcitarsis (Broun, 1880), new combination Fig. 50, 93, 153 Anchomenus (Platynus) sulcitarsis Broun, 1880: 27. Holotype: “one mutilated specimen at Parua, near Whangarei Harbour” [ND] (BMNH, could not be located). Anchomenus sulcitarsis: Hutton 1904: 144. Agonum (Platynus) sulcitarse: Csiki 1931: 853. Agonum sulcitarse: Watt 1971: 25. Ctenognathus sulcitarsis: Larochelle and Larivière 2007: 81, 111. Description. Body length 12.8–14.0 mm. Head, pronotum, and elytra black; abdomen piceous black; antennae and palpi reddish; legs dark rufous. Microsculpture isodiametric on head, very transverse (with microlines) on pronotum, and isodiametric on elytra. Iridescence present on pronotum. Very shiny. Head. Moderately wide. Mandibles moderately long, moderately curved anteriorly. Labrum moderately transverse, subtruncate anteriorly. Tempora moderately inflated and short. Submentum with six setae. Thorax. Pronotum moderately convex, obsoletely punctate across base, obsoletely wrinkled on disc, narrow, cordate, widest about middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead complete; sides moderately rounded, not sinuate posteriorly; lateral beads incomplete, obsolete basally; lateral depressions wide, incomplete near apex; posterolateral angles obtuse; laterobasal foveae very deep, narrow, prolonged forward; posterior bead incomplete, absent medially; base subtruncate. Apex of prosternum pubescent. Legs. Moderately long. Metacoxae with three posteroventral setae. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, shallowly bisulcate laterally. Metatarsomeres 4 moderately asymmetrical apically. Elytra. Slightly convex, sloping down toward apex, oblong. Basal margin complete, reaching about scutellum. Shoulders obtuse. Sides slightly rounded. Scutellar setiferous pore invisible. Scutellar striole impunctate. Striae deep, impunctate. Intervals depressed. Umbilicate series with 17–18 setiferous punctures. Subapical sinuations feeble. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 93): base strongly convex dorsally, with basal lobe very wide; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area narrow and moderately long; internal sac without scale-like sclerites; apex subtriangular, moderately convex dorsally, slightly concave ventrally, with extreme tip moderately wide and long. Dorsal view: moderately wide, symmetrical (ostium of membranous area dorsal); basal bulb close to membranous area. Material examined. 224 specimens (AMNZ, BMNH, CMNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 153). North Island: AK, CL, ND, WO. 30 · April 30, 2021 Larochelle and Larivière Key to species of Ctenognathus 1. Pronotum with a single setiferous puncture on each side (anteriorly) .............................2 — Pronotum with two setiferous punctures on each side (anteriorly and posteriorly) .................9 2(1). Mesoand metatarsomeres 1–4 (Fig. 27) not carinate dorsally ...................................3 — Mesoand metatarsomeres 1–4 (Fig. 28) tricarinate dorsally ....................................6 3(2). Antennae entirely rufopiceous; abdomen black . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .4 — Antennae bicolored; abdomen rufopiceous ...................................................5 4(3). Tempora not inflated; pronotum not sinuate posteriorly, sides moderately rounded, posterolateral angles obtusely rounded, base truncate, posterior bead present, complete; apices of elytra obtuse; Fig. 55. [Body length 12.0–13.9 mm; North Island (ND to TK–HB)] ................................. ................................................Ctenognathus novaezelandiae (Fairmaire) — Tempora moderately inflated (about two-thirds as long as eyes); pronotum moderately sinuate posteriorly, sides strongly rounded, posterolateral angles slightly acute, base convex, posterior bead absent; apices of elytra rounded; Fig. 56. [Body length 12.0–13.2 mm; southern North Island (WN)] ..... ..........................................................Ctenognathus elevatus (White) 5(3). Antennal segments 1–4 rufopiceous, 5–11 reddish; mandibles very long, slightly curved anteriorly; pronotum with lateral depressions narrow, slightly widened from apex to base, posterolateral angles obtuse; metatarsomeres 5 glabrous ventrally; Fig. 57. [Body length 11.4–12.8 mm; Three Kings Islands (TH)] .................. Ctenognathus garnerae Larochelle and Larivière, new species — Antennal segments 1 rufopiceous, 2–11 reddish; mandibles moderately long and curved anteriorly; pronotum with lateral depressions wide throughout, posterolateral angles rectangular; metatarsomeres 5 setose ventrally (with long setae); Fig. 58. [Body length 11.6–13.2 mm; northern North Island (ND, top of Aupouri Peninsula)] ....Ctenognathus tepaki Larochelle and Larivière, new species 6(2). Head, pronotum, and elytra coal black; femora rufopiceous; elytra without visible scutellar setiferous pore, microsculpture very transverse (with microlines); terminal segment of palpi obtuse apically (Fig. 10) ............................................................................ 7 — Head and pronotum pure black, elytra piceous black; femora reddish; elytra with visible scutellar setiferous pore, microsculpture isodiametric; terminal segment of palpi truncate apically (Fig. 11) .... 8 7(6). Pronotum widest before middle, slightly wrinkled on disc, sides strongly sinuate posteriorly, posterolateral angles rectangular; interval 3 of elytra with a minute subapical setiferous puncture; Fig. 59. [Body length 12.7–13.8 mm; eastern central North Island (BP, GB, HB)] ....................... .............................. Ctenognathus urewera Larochelle and Larivière, new species — Pronotum widest about middle, deeply wrinkled on disc, sides not sinuate posteriorly, posterolateral angles acute and projected laterally; interval 3 of elytra without setiferous punctures; Fig. 60. [Body length 11.3–13.8 mm; northern North Island (ND to WO)] .................................. ..................................................Ctenognathus cardiophorus (Chaudoir) 8(6). Pronotum subquadrate, moderately convex, sides moderately sinuate posteriorly, posterolateral angles rectangular, moderately long; interval 3 of elytra without setiferous punctures; Fig. 61. [Body length 12.7–12.9 mm; North Island (AK–CL to WN)] ................. Ctenognathus adamsi (Broun) — Pronotum elongate, slightly convex, sides strongly sinuate posteriorly, posterolateral angles acute (projected laterally), very long; interval 3 of elytra with a large (foveate) subapical setiferous puncture; Fig. 62. [Body length 11.0–13.6 mm; southern North Island (WI to WN–WA) and northern South Island (SD, NN, MB)] . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . Ctenognathus pictonensis Sharp 9(1). Femora dark, pure black, piceous black or dark rufous ....................................... 10 — Femora paler, yellow or rufotestaceous ..................................................... 15 10(9). Scutellar setiferous pore of elytra invisible; legs short; Fig. 63. [Rather small, body length 7.0–9.8 mm; terminal segment of palpi truncate apically; southern South Island (SL)] ....................... .............................. Ctenognathus tawanui Larochelle and Larivière, new species Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 31 — Scutellar setiferous pore of elytra visible; legs moderately long or very long ..................... 11 11(10). Pronotum strongly cordate; elytra subovate, strongly convex; Fig. 64. [Body length 11.9–12.4 mm; southwestern South Island (southern WD)] .................................................... ............................ Ctenognathus perumalae Larochelle and Larivière, new species — Pronotum moderately cordate, subquadrate, or subrectangular; elytra oblong, slightly convex or subdepressed ............................................................................ 12 12(11). Antennal segments 1–3(4) black, (3)4–11 dark rufous ........................................ 13 — Antennae entirely dark rufous ............................................................ 14 13(12). Femora and tibiae black; pronotum moderately cordate, base emarginate; elytra subdepressed, microsculpture isodiametric, striae finely punctate; Fig. 65. [Body length 8.5–11.4 mm; southwestern South Island (southern FD)] ............ Ctenognathus takahe Larochelle and Larivière, new species — Femora and tibiae dark rufous; pronotum subrectangular, base subtruncate; elytra slightly convex, microsculpture moderately transverse, striae impunctate; Fig. 66. [Body length 10.1–10.6 mm; northern South Island (NN, BR, MB, KA)] ............... Ctenognathus arnaudensis (Broun) 14(12). Pronotum unwrinkled, moderately cordate, laterobasal foveae moderately wide (not meeting toward pronotal midline); elytra very shiny, slightly convex, interval 3 with three or four large (foveate) setiferous punctures; Fig. 67. [Body length 12.6–13.3 mm; western South Island (BR, WD, OL)] .. ......................................Ctenognathus marieclaudiae Larochelle, new species — Pronotum wrinkled on disc and across base, subquadrate, laterobasal foveae very wide (meeting toward pronotal midline); elytra dull, subdepressed, interval 3 with three shallow, minute setiferous punctures; Fig. 68. [Smaller, body length 8.5–11.0 mm; southern South Island (MC to FD–SL) and Stewart Island] ................................................... Ctenognathus edwardsii (Bates) 15(9). Mesoand metatarsomeres 1–4 (Fig. 27) not carinate dorsally; Fig. 69. [Body length 9.1–10.5 mm; northeastern South Island (MB, KA)] ..................................................... ..............................Ctenognathus kaikoura Larochelle and Larivière, new species — Mesoand metatarsomeres 1–4 (Fig. 28) tricarinate dorsally ...................................16 16(15). Sternum VII (last visible sternum of abdomen; Fig. 35): male with two long apical ambulatory setae, female with four long apical ambulatory setae ............................................17 — Sternum VII (Fig. 36): male with four long apical ambulatory setae, female with eight to ten long apical ambulatory setae .....................................................................20 17(16). Posterolateral angles of pronotum acute and projected laterally; terminal segment of palpi obtuse apically (Fig. 10) ..............................................................................18 — Posterolateral angles of pronotum rectangular; terminal segment of palpi truncate apically (Fig. 11) ...19 18(17). Femora and tibiae pale yellow; pronotum with anterolateral angles poorly developed, posterior bead incomplete (obsolete medially), base emarginate; legs moderately long; elytra very convex; Fig. 70. [Body length 10.2–11.1 mm; southern North Island (WN, WA)] ............................ ......................................................Ctenognathus sandageri (Broun) — Femora dark yellow, tibiae reddish; pronotum with anterolateral angles well developed, posterior bead complete, base subtruncate; legs very long; elytra moderately convex; Fig. 71. [Body length 9.5–11.6 mm; western South Island (NN–SD to WD)] ................... Ctenognathus helmsi (Sharp) 19(17). Elytra ovate; pronotum widest about middle, sides strongly rounded; Fig. 72. [Body length 10.0–11.4 mm; North Island (WO–BP to WN)] ....................Ctenognathus xanthomelus (Broun) — Elytra subovate; pronotum widest before middle, sides moderately rounded; Fig. 73. [Body length 10.5– 12.0 mm; southern North Island (TO to WN)] .............Ctenognathus intermedius (Broun) 20(16). Femora and tibiae pale yellow ............................................................ 21 — Femora dark yellow or reddish; tibiae reddish ................................................23 21(20). Elytra shiny; posterolateral angles of pronotum slightly obtuse; mentum tooth subtruncate apically; submentum with ten long setae; Fig. 75. [Body length 10.8–11.2 mm; pronotum slightly cordate; elytra 32 · April 30, 2021 Larochelle and Larivière slightly convex, striae finely punctate; South Island (NN–BR to NC–MK)] ..................... ........................................................Ctenognathus colensonis (White) — Elytra dull; posterolateral angles of pronotum rectangular; mentum tooth emarginate apically; submentum with six long setae ............................................................... 22 22(21). Pronotum slightly cordate; elytra subdepressed, striae finely punctate; eyes strongly rounded; terminal segment of palpi obtuse apically; Fig. 74. [Body length 10.3–12.7 mm; southern South Island (OL– DN to FD–SL)] Ctenognathus otagoensis (Bates) — Pronotum subquadrate; elytra moderately convex, striae impunctate; eyes moderately rounded; terminal segment of palpi truncate apically; Fig. 76. [Body length 10.2–11.5 mm; Stewart Island] .......... .................................Ctenognathus earlyi Larochelle and Larivière, new species 23(20). Antennal segments 1–3 rufotestaceous, 4–11 dark piceous; femora rufotestaceous; sides of pronotum not sinuate posteriorly, lateral depressions wide throughout, laterobasal foveae very wide (almost meeting toward pronotal midline); elytra with sides slightly rounded, interval 3 with three very deep, large (foveate) setiferous punctures; Fig. 77. [Body length 11.2–12.9 mm; western South Island (BR–SL)] .....................Ctenognathus davidsoni Larochelle and Larivière, new species — Antennae entirely dark rufous or rufotestaceous; femora dark yellow; sides of pronotum moderately sinuate posteriorly, lateral depressions narrow anteriorly and widened posteriorly, laterobasal foveae moderately wide (well separated from each other); elytra with sides moderately rounded, interval 3 with three shallow, minute setiferous punctures ......................................... 24 24(23). Antennae dark rufous; terminal segment of palpi obtuse apically; eyes strongly convex; pronotum obsoletely wrinkled on disc, moderately cordate, posterolateral angles rectangular; elytra oblong, sides moderately rounded, striae deep and impunctate; Fig. 78. [Body length 10.5–13.00 mm; southeastern South Island (MK–SL)] .................................Ctenognathus oreobius (Broun) — Antennae rufotestaceous; terminal segment of palpi truncate apically; eyes moderately convex; pronotum deeply wrinkled on disc, slightly cordate, posterolateral angles acute and slightly projected laterally; elytra subovate, sides strongly rounded, striae shallow and finely punctate; Fig. 79. [Body length 11.1–12.3 mm; Banks Peninsula (MC)] ............................................. ................................ Ctenognathus hoarei Larochelle and Larivière, new species Ctenognathus novaezelandiae (Fairmaire, 1843) Fig. 55, 98, 138 Anchomenus Novae-Zelandiae Fairmaire, 1843: 12. Holotype: one specimen from “Nouvelle-Zélande” [= New Zealand] (MNHN, Fairmaire Collection, could not be located). Anchomenus (Ctenognathus) novae zeelandiae [sic]: White 1846: 3. Ctenognathus novae zelandiae: Chenu 1851: 139. Dicrochile ovicollis Motschulsky, 1865: 316. Holotype: one specimen from “Nouvelle-Zélande” [= New Zealand]. Synonymized by Kuschel (1990: 76). Anchomenus punctulatus Broun, 1877: 371. Holotype: “Habitat: Auckland” (BMNH, could not be located). New synonym Colpodes neozelandicus Chaudoir, 1878: 294. Lectotype (here designated): male (MNHN) labeled “baie des isles [= ND, Bay of Islands] (hand-written) / ExMusaeo Chaudoir (typed) / neozelandicus Chaud. Nouv. Zélande Baie des Isles Coll. Reiche (hand-written) / LECTOTYPE [male symbol] Colpodes neozelandicus Chaudoir, 1878 designated by Larochelle & Larivière, 2021 (red label; typed).” New synonym Anchomenus montivagus Broun, 1880: 22. Holotype: one specimen “near Whangarei Heads” [ND] (BMNH, could not be located). New synonym Anchomenus politulus Broun, 1880: 22. Holotype: female (BMNH) labeled “Type (circular red-bordered label; typed) / 47. (hand-written) / [ND] [Mt] Manaia 1876 (hand-written) / New Zealand. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; typed) / Ctenognathus politulus. (hand-written).” New synonym Anchomenus suborbithorax Broun, 1880: 24. Holotype: male (BMNH) labeled “Type (circular red-bordered label; typed) / 51. [male symbol] (hand-written) / [ND] [Mt] Manaia 1877 (hand-written) / New Zeal. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; typed) / Ctenognathus suborbithorax (hand-written).” New synonym Anchomenus perrugithorax Broun, 1880: 24. Holotype: male (BMNH) labeled “Type (circular red-bordered label; typed) / 52. [male symbol] (hand-written) / New Zeal. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 33 typed) / [ND] Mt Manaia 1878 (hand-written) / Ctenognathus perrugithorax. (hand-written).” New synonym Ctenognathus latipennis Sharp, 1886: 363. Synonymized with Anchomenus montivagus Broun, 1882 by Csiki (1931: 744). Ctenognathus politulus: Broun 1893: 986. Ctenognathus suborbithorax: Broun 1893: 986. Ctenognathus perrugithorax: Broun 1893: 986. Ctenognathus neozealandicus [sic]: Hutton 1904: 144 Anchomenus suborbithorax: Walker 1904: 115. Ctenognathus suborbithorax: Hutton 1904: 144. Colpodes neozelandicus: Csiki 1931: 758. Agonum (Anchomenus) punctatum [sic]: Csiki 1931: 865. Ctenognathus neozelandius [sic]: Hudson 1934: 174. Agonum novaezealandicum [sic]: Moeed and Meads 1985: 22. Ctenognathus novaezelandiae: Kuschel 1990: 24, 40. Ctenognathus montivagus: Larochelle and Larivière 2001: 135. Ctenognathus punctulatus: Larochelle and Larivière 2007: 81, 111. Description. Body length 12.0–13.9 mm. Head, pronotum, elytra, and abdomen black; antennae, palpi, and legs rufopiceous. Microsculpture very transverse (with microlines) on head and pronotum, isodiametric on elytra. Iridescence present on head and pronotum. Very shiny on head and pronotum, moderately shiny on elytra. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, moderately emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth entire, rounded apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc and across base, wide, slightly cordate, widest before middle; apex strongly emarginate; anterolateral angles well developed, broadly rounded; anterior bead complete; sides moderately rounded, not sinuate posteriorly; lateral beads complete; lateral depressions wide, more so posteriorly; a single setiferous puncture on each side (anteriorly); posterolateral angles obtusely rounded; laterobasal foveae moderately deep and wide, prolonged forward; posterior bead complete; base truncate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 poorly developed, not carinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 setose ventrally (with long setae). Elytra. Slightly convex, not sloping down toward apex, oblong, widest about middle. Shoulders slightly rounded. Sides moderately rounded. Scutellar setiferous pore invisible. Scutellar striole short, impunctate. Striae deep, impunctate. Intervals moderately convex; interval 3 with a single very deep, large (foveate) subapical setiferous puncture. Umbilicate series with 19–20 setiferous punctures. Subapical sinuations moderately strong. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with nine or ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 98): strongly arcuate, slender; base moderately convex dorsally, with basal lobe very wide; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area strongly widened in apical half and moderately long; apex moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and very long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 1421 specimens (AMNZ, BMNH, CMNH, JNNZ, LUNZ, MNHN, MONZ, NZAC). Geographic distribution (Fig. 138). North Island: AK, BP, CL, HB, ND, TK, WO. Ecology. Coastal lowland. Epigean, fossorial, arboreal. Sand dunes and adjacent forests (broadleaf, podocarp), shrublands, and tree plantations (pine). Mostly open ground; dry soil. Nocturnal; hides during the day in burrows at the base of plants (Muehlenbeckia, grass), in leaf litter, under logs, fallen trees and branches, or stones. Gregarious. Larval habitat: leaf litter and rotten logs. Biology. Seasonality: Throughout the year; numbers decreasing from mid-winter onward. Tenerals: Spring. Young adults mostly inactive over summer. An autumn-winter breeder. Egg stage: 9 days. Larval development: 34 · April 30, 2021 Larochelle and Larivière 82.2 days (autumn and winter). Pupal stage: 11.8 days. Adult lifespan: one year. Predacious. Food: Small arthropods, e.g., insects. Predators: Rats. Occasionally infested with mites. Dispersal power. Subapterous (incapable of flight). Moderate runner. Occasional burrower. Frequent climber (on trees). Collecting techniques. Examining burrows at the base of Muehlenbeckia plants, pitfall trapping; sifting leaf litter; turning logs, fallen branches, and stones; pan trapping (yellow pan traps). References. Larochelle and Larivière 2001: 131, 135–136 (as “Anchomenus” punctulatus, Ctenognathus montivagus, C. neozelandicus, C. novaezelandiae, C. perrugithorax, C. politulus, C. suborbithorax; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus punctulatus; list); Philip and Burgess, 2008a: 35–39 (life history, rearing), 2008b: 41–46 (behavior, ecology), Larochelle and Larivière 2016: 37 (as Ctenognathus montivagus, C. neozelandicus, C. novaezelandiae, C. perrugithorax, C. politulus, Ctenognathus punctulatus, C. suborbithorax; list). Remarks. The holotype of Anchomenus Novae-Zelandiae could not be located in MNHN where Fairmaire’s specimens can usually be found. However, there is no ambiguity regarding the identity of this taxon; the original description can only apply to the species as currently understood. In addition to diagnostic characters of the male genitalia, Ctenognathus novaezelandiae has the following distinguishing features: tempora not inflated; pronotum not sinuate posteriorly, sides moderately rounded, posterolateral angles obtusely rounded, base truncate, posterior bead present, complete; elytral apices obtuse. This species occurs in coastal dunes and forests on the North Island (ND to TK–HB). The holotypes of Anchomenus punctulatus Broun and Anchomenus montivagus Broun could not be located in BMNH where Broun’s specimens can usually be found. However, there is no ambiguity regarding the identity of these taxa; their original descriptions can only apply to Ctenognathus novaezelandiae as currently understood. Chaudoir described Colpodes neozelandicus from three specimens, one of which (a male) could be seen (MNHN) and is here designated as lectotype in order to preserve stability of nomenclature in the future. Examination of this specimen revealed it to be conspecific with C. novaezelandiae. Finally, examination of Broun’s types of Anchomenus politulus, A. suborbithorax and A. perrugithorax also revealed them to be conspecific with Ctenognathus novaezelandiae. Ctenognathus elevatus (White, 1846), reinstated Fig. 56, 99, 131 Anchomenus elevatus White, 1846: 3. Holotype: male (BMNH) labeled “Type H.T. (circular red-bordered label; typed) / New Zealand (hand-written) / Anchomenus elevatus Zool. Erebus & Terror (hand-written) / Anchomenus elevatus. White (hand-written).” Erroneously synonymized with Anchomenus novaezelandiae Fairmaire, 1843 by Sharp (1884: 297). Reinstated as full species Ctenognathus actochares Broun, 1894: 307. Holotype: male (BMNH) labeled “Type (circular red-bordered label; typed) / 2660 (hand-written) / [WN] Wellington (hand-written) / New Zealand. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; typed) / Ctenognathus actochares (hand-written).” New synonym Ctenognathus (Anchomenus) actochares: Walker 1904: 76. Ctenognathus actochares: Hudson 1923: 356. Description. Body length 12.0–13.2 mm. Head, pronotum, elytra, and abdomen black; antennae, palpi, and legs rufopiceous. Microsculpture very transverse (with microlines) on head and pronotum, isodiametric on elytra. Iridescence present on head and pronotum. Very shiny on head and pronotum, moderately shiny on elytra. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about two-thirds as long as eyes). Neck constriction shallow dorsolaterally. Mentum: medial tooth entire, rounded apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, obsoletely punctate across base, obsoletely wrinkled on disc and across base, wide, moderately cordate, widest before middle; apex strongly emarginate; anterolateral angles well developed, broadly rounded; anterior bead complete; sides strongly rounded, moderately sinuate posteriorly; lateral beads complete; lateral depressions wide, slightly widened near posterolateral angles; a single setiferous puncture on each side (anteriorly); posterolateral angles rectangular or Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 35 slightly acute (projected laterally); laterobasal foveae moderately deep and wide, prolonged forward; posterior bead absent; base convex. Legs. Moderately long. Metafemora with three posteroventral setae. Mesoand metatarsomeres 1–4 poorly developed, not carinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 setose ventrally (with long setae). Elytra. Slightly convex, not sloping down toward apex, oblong, widest about middle. Shoulders slightly rounded. Sides moderately rounded. Scutellar setiferous pore invisible. Scutellar striole short, impunctate. Striae deep, impunctate. Intervals moderately convex; interval 3 with a single very deep, large (foveate) subapical setiferous puncture. Umbilicate series with 19–20 setiferous punctures. Subapical sinuations moderately strong. Apices rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with nine or ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 99): strongly arcuate; base strongly convex and biconcave dorsally, with basal lobe narrow; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area moderately wide and moderately long; apex strongly convex dorsally, slightly concave ventrally, with extreme tip narrow and very long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 95 specimens (AMNZ, BMNH, CMNH, JNNZ, MONZ, NZAC). Geographic distribution (Fig. 131). North Island: WN. Ecology. Coastal lowland. Epigean, fossorial. Sand dunes and adjacent scrublands or pine forests. Mostly open ground; dry soil. Nocturnal; hides during the day in burrows at the base of plants (Muehlenbeckia), or under stones and logs. Gregarious. Biology. Seasonality: Throughout the year. Tenerals: October–November. Predacious (based on mouthpart morphology). Predators: Spiders. Occasionally infested with mites and fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Examining burrows at the base of Muehlenbeckia plants; turning stones and logs. References. Larochelle and Larivière 2001: 133, 135 (as junior synonym of Ctenognathus novaezelandiae, as C. actochares; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 111 (as C. actochares; list); 2016: 36 (as C. actochares; list). Remarks. Anchomenus elevatus was erroneously synonymized with Ctenognathus novaezelandiae by Sharp (1884). However, the holotype was seen and there is no ambiguity regarding the identity of this taxon which is here resurrected from synonymy. In addition to diagnostic characters of the male genitalia, C. elevatus has the following distinguishing features: tempora moderately inflated (about two-thirds as long as eyes); pronotum moderately sinuate posteriorly, sides strongly rounded, posterolateral angles slightly acute, base convex, posterior bead absent; elytral apices rounded. The two species are allopatric: C. elevatus is restricted to southern areas of the North Island (WN), while C. novaezelandiae occurs in more northern areas of the North Island (ND to TK–HB). Examination of the type of Ctenognathus actochares Broun, revealed it to be conspecific with Ctenognathus elevatus. Anchomenus elevatus Bates, 1874 is a nomen nudum (Larochelle and Larivière 2001: 135, 178). Ctenognathus garnerae Larochelle and Larivière, new species Fig. 57, 100, 132 Ctenognathus garnerae Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “[NEW ZEALAND TH] Three Kings Is Great I. Nov. 70 NZ. Ent. Div. Exp. (typed) / Castaway Camp (typed) / G. Kuschel (typed) / ex. crevices in coastal cliffs. (hand-written) / HOLOTYPE [male symbol] Ctenognathus garnerae Larochelle & Larivière, 2021 (red label; typed).” Paratypes: one female (NZAC) and one male (MONZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 11.4–12.8 mm. Head, pronotum, and elytra black; abdomen rufopiceous; antennal segments 1–4 rufopiceous, 5–11 reddish; palpi reddish; femora and tibiae rufopiceous; tarsi reddish. Microsculpture very transverse (with microlines) on head and pronotum, isodiametric on elytra. Iridescence present on pronotum. Very shiny on head and pronotum, moderately shiny on elytra. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles very long, slightly curved anteriorly. Labrum strongly 36 · April 30, 2021 Larochelle and Larivière transverse, moderately emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth entire, rounded apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc and across apex, wide, moderately cordate, widest before middle; apex strongly emarginate; anterolateral angles well developed, broadly rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, moderately sinuate posteriorly; lateral beads incomplete, obsolete anteriorly; lateral depressions narrow, slightly widened from apex to base; a single setiferous puncture on each side (anteriorly); posterolateral angles obtuse; laterobasal foveae moderately deep and wide, prolonged forward; posterior bead obsolete; base emarginate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 poorly developed, not carinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Slightly convex, not sloping down toward apex, oblong, widest about middle. Shoulders slightly rounded. Sides moderately rounded. Scutellar setiferous pore invisible. Scutellar striole short, impunctate. Striae deep, impunctate. Intervals moderately convex; interval 3 with a single very deep, large (foveate) subapical setiferous puncture. Umbilicate series with 21 setiferous punctures. Subapical sinuations moderately strong. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 100): strongly arcuate, moderately wide; base strongly convex dorsally, with basal lobe moderately wide; middle moderately convex, slightly concave in apical half dorsally, strongly concave ventrally, with dorsal membranous area moderately wide and moderately long; apex strongly convex dorsally, slightly concave ventrally, with extreme tip moderately wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 19 specimens (MONZ, NZAC). Geographic distribution (Fig. 132). Offshore Islands: TH–Great Island, Castaway Camp. Ecology. Coastal lowland. Epigean, fossorial. Sea cliffs and adjacent forests. Open or shaded ground; dry soil. Nocturnal; hides during the day in rock crevices. Gregarious. Biology. Seasonality: November. Tenerals: September. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Examining rock crevices; pan trapping (yellow pan traps). Remarks. This species is named after our friend and colleague Beulah Garner (The Natural History Museum, London (BMNH)) for facilitating our study of the type material of Carabidae, especially the Broun Collection. Ctenognathus garnerae is morphologically close to C. tepaki. In addition to diagnostic characters of the male genitalia, C. garnerae has the following distinguishing features: antennal segments 1–4 rufopiceous, 5–11 reddish; mandibles very long, slightly curved anteriorly; pronotum with lateral depressions narrow and slightly widened from apex to base, posterolateral angles obtuse; metatarsomeres 5 glabrous ventrally. The two species are allopatric: C. garnerae is restricted to the Three Kings Islands (TH), while C. tepaki occurs in northernmost areas of the North Island (ND, top of Aupouri Peninsula). Ctenognathus tepaki Larochelle and Larivière, new species Fig. 58, 101, 146 Ctenognathus tepaki Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “[NEW ZEALAND ND] Spirits Bay, N.Z. (typed) under stone. (hand-written) 20–28:VIII:1957 J.C. Watt. (typed) / J.C. Watt Collection Ent. Div. DSIR, 1966. (typed) / HOLOTYPE [male symbol] Ctenognathus tepaki Larochelle & Larivière, 2021 (red label; typed).” Paratypes: one female (NZAC) from the same locality as the holotype and one male (AMNZ) from Cape Maria van Diemen, Motuopao Island (ND), bearing blue paratype labels. Description. Body length 11.6–13.2 mm. Head, pronotum, and elytra black; abdomen rufopiceous; antennae reddish (scape rufopiceous); palpi reddish; femora and tibiae rufopiceous; tarsi reddish. Microsculpture very transverse (with microlines) on head and pronotum, isodiametric on elytra. Iridescence present on head and Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 37 pronotum. Very shiny on head and pronotum, moderately shiny on elytra. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, moderately emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth entire, rounded apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc and across base, wide, moderately cordate, widest before middle; apex strongly emarginate; anterolateral angles well developed, broadly rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, moderately sinuate posteriorly; lateral beads incomplete, obsolete anteriorly; lateral depressions wide; a single setiferous puncture on each side (anteriorly); posterolateral angles rectangular; laterobasal foveae moderately deep and wide, prolonged forward; posterior bead incomplete, obsolete medially and near posterolateral angles; base emarginate. Legs. Moderately long. Metafemora with three posteroventral setae. Mesoand metatarsomeres 1–4 poorly developed, not carinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 setose ventrally (with long setae). Elytra. Slightly convex, not sloping down toward apex, oblong, widest about middle. Shoulders slightly rounded. Sides moderately rounded. Scutellar setiferous pore invisible. Scutellar striole short, impunctate. Striae deep, impunctate. Intervals moderately convex; interval 3 with a single very deep, large (foveate) subapical setiferous puncture. Umbilicate series with 21–24 setiferous punctures. Subapical sinuations moderately strong. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with eight to ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 101): strongly arcuate, slender; base slightly convex, with basal lobe very wide; middle strongly biconvex dorsally, strongly concave ventrally, with dorsal membranous area moderately wide in apical half and moderately long; apex slender, moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and very long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 74 specimens (AMNZ, CMNZ, MONZ, NZAC). Geographic distribution (Fig. 146). North Island: ND (Te Paki area)– Huka Creek [=Stream]. Matapia Island. Maungapiko [Hill]. Motuopao Island. Murimotu Island. Ngaroku Stream. Spirits Bay. Unuwhao. Ecology. Coastland lowland. Epigean, fossorial. Sand dunes and adjacent forests. Shaded ground; dry soil. Nocturnal; hides during the day in burrows at the base of plants (Muehlenbeckia) or under stones. Biology. Seasonality: October–January, August. Tenerals: November. Predacious (based on mouthpart morphology). Occasionally infested with mites and fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Examining burrows at the base of Muehlenbeckia plants; turning stones. Remarks. The species is named after the Te Paki area (ND, Aupouri Peninsula) which includes the type locality Spirits Bay, and is applied as a noun in apposition. Ctenognathus tepaki is morphologically close to C. garnerae. In addition to diagnostic characters of the male genitalia, C. tepaki has the following distinguishing features: antennal segments 1 rufopiceous, 2–11 reddish; mandibles moderately long and curved anteriorly; pronotum with lateral depressions wide, posterolateral angles rectangular; metatarsomeres 5 setose ventrally (with long setae). The two species are allopatric: C. tepaki is found in northern North Island (ND, top of Aupouri Peninsula), while C. garnerae is restricted to the Three Kings Islands (TH). Ctenognathus urewera Larochelle and Larivière, new species Fig. 59, 102, 147 Ctenognathus urewera Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “NEW ZEALAND GB Urewera NP, Huiarau Summit 923m 14–18.II.1994 Larivière, Larochelle (typed) / Nothofagus + Horopito + tree fern for. [= forest], wet slopes. Pittraps. (typed) / HOLOTYPE [male symbol] Ctenognathus urewera Larochelle & Larivière, 2021 (red label; typed).” Paratypes: two females (MONZ, NZAC) from the same locality as the holotype, bearing blue paratype labels. 38 · April 30, 2021 Larochelle and Larivière Description. Body length 12.7–13.8 mm. Head, pronotum, and elytra coal black; abdomen blackish; antennae, palpi, and femora rufopiceous; tibiae and tarsi reddish. Microsculpture very transverse (with microlines) on head, pronotum, and elytra. Iridescence present on head and pronotum. Very shiny. Forebody narrow in comparison to elytra. Head. Narrow. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, moderately emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth truncate apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, obsoletely punctate across apex and base, slightly wrinkled on disc, narrow, strongly cordate, widest before middle; apex strongly emarginate; anterolateral angles well developed, angulate; anterior bead complete, less impressed medially; sides strongly rounded and sinuate posteriorly; lateral beads complete; lateral depressions narrow; a single setiferous puncture on each side (anteriorly); posterolateral angles rectangular; laterobasal foveae moderately deep, rather narrow, prolonged forward; posterior bead complete, less impressed medially; base emarginate. Legs. Very long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Slightly convex, not sloping down toward apex, subovate, widest about middle. Shoulders moderately rounded. Sides strongly rounded. Scutellar setiferous pore invisible. Scutellar striole short or moderately long, impunctate. Striae obsolete, impunctate. Intervals slightly convex; interval 3 with a single shallow, minute subapical setiferous puncture. Umbilicate series with 17–18 setiferous punctures. Subapical sinuations feeble. Apices rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 102): strongly arcuate; base strongly convex dorsally, with basal lobe very wide; middle moderately convex in basal half and straight in apical half dorsally, slightly concave ventrally, with dorsal membranous area very wide and very long; apex subtriangular, straight dorsally and ventrally, with extreme tip narrow and short. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 811 specimens (JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 147). North Island: BP, GB, HB. Ecology. Lowland, montane. Epigean, fossorial, arboreal. Wet forests (beech, broadleaf) and tree plantations (pine). Associated with stream, seepages, and mud flats. Shaded ground; wet, soaked soil. Nocturnal; hides during the day under logs, fallen branches, and stones. Gregarious. Biology. Seasonality: September–January, March–April. Tenerals: December–February. Predacious (based on mouthpart morphology). Occasionally infested with mites and fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Occasional climber (on trees). Collecting techniques. Pitfall trapping; turning logs, fallen branches, and stones. Remarks. The species is named after the Urewera National Park (GB) which includes the type locality Huiarau Summit, and is applied as a noun in apposition. Ctenognathus urewera is morphologically close to C. cardiophorus. In addition to diagnostic characters of the male genitalia, C. urewera has the following distinguishing features: pronotum widest before middle, slightly wrinkled on disc, sides strongly sinuate posteriorly, posterolateral angles rectangular; interval 3 of elytra with a minute subapical setiferous puncture. The two species are allopatric: C. urewera occurs in eastern central areas of the North Island (BP, GB, HB), while C. cardiophorus is found in more northern areas (ND–WO). Ctenognathus cardiophorus (Chaudoir, 1878) Fig. 60, 103, 126 Colpodes cardiophorus Chaudoir, 1878: 305. Holotype: female (MNHN) labeled “ExMusaeo Chaudoir (typed) / cardiophorus Chaud. Nouv. Zélande C. Laferté (hand-written).” Anchomenus parabilis Broun, 1880: 20. Holotype: “Whangarei Heads” [ND] (BMNH could not be located). New synonym Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 39 Ctenognathus parabilis: Broun 1893: 986. Ctenognathus cardiophorus: Hutton 1904: 144. Colpodes cardiophorus: Csiki 1931: 749. Ctenognathus cardiophorus: Hudson 1934: 174. Description. Body length 11.3–13.8 mm. Head, pronotum, and elytra coal black; abdomen blackish; antennae and palpi reddish; femora rufopiceous; tibiae and tarsi reddish. Microsculpture very transverse (with microlines) on head, pronotum, and elytra. Iridescence absent. Moderately shiny. Forebody narrow in comparison to elytra. Head. Narrow. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth truncate apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, deeply wrinkled on disc, narrow, strongly cordate, widest about middle; apex slightly emarginate; anterolateral angles well developed, angulate; anterior bead complete; sides strongly rounded, not sinuate posteriorly; lateral beads complete; lateral depressions wide, more so posteriorly; a single setiferous puncture on each side (anteriorly); posterolateral angles acute, projected laterally; laterobasal foveae unusually depressed, shallow, moderately wide, prolonged forward; posterior bead complete, less impressed near posterolateral angles; base emarginate. Legs. Very long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed and asymmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Moderately convex, not sloping down toward apex, subovate, widest about middle. Shoulders moderately rounded. Sides strongly rounded. Scutellar setiferous pore invisible. Scutellar striole short, impunctate. Striae obsolete, impunctate. Intervals moderately convex; interval 3 without setiferous punctures. Umbilicate series with 18–21 setiferous punctures. Subapical sinuations moderately strong. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with eight long apical ambulatory setae. Aedeagus. Lateral view (Fig. 103): strongly arcuate; base strongly convex, with basal lobe very wide; middle strongly convex dorsally, moderately concave ventrally, with dorsal membranous area biconvex, strongly widened in apical half and very long; apex moderately convex dorsally, rather straight ventrally, with extreme tip moderately wide and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 519 specimens (AMNZ, BMNH, CMNH, LUNZ, MNHN, MONZ, NZAC). Geographic distribution (Fig. 126). North Island: AK, CL, ND, WO. Ecology. Lowland. Epigean, fossorial. Wet forests (broadleaf, podocarp) and tree plantations (pine). Associated with streams and slopes. Shaded ground; wet, soaked soil. Nocturnal; hides during the day under logs, fallen branches, and stones. Gregarious. Biology. Seasonality: Throughout the year. Tenerals: November–January. Predacious (based on mouthpart morphology). Predators: Spiders. Occasionally infested with mites. Dispersal power. Subapterous (incapable of flight). Swift runner. Collecting techniques. Pitfall trapping; turning logs, fallen branches, and stones. References. Larochelle and Larivière 2001: 134, 136 (as Ctenognathus cardiophorus, C. parabilis, catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 111 (as C. cardiophorus, C. parabilis; list), 2016: 36–37 (as C. cardiophorus, C. parabilis; list). Remarks. The holotype of Anchomenus parabilis could not be located in BMNH where Broun’s specimens can usually be found. However, there is no ambiguity regarding the identity of this taxon. Furthermore, a non-type male specimen from the Broun Collection (BMNH) was examined. This specimen is from Hunua-Wairoa [= Wairoa, Hunua Ranges] (AK) and its morphology corresponds to Broun’s original description of Ctenognathus cardiophorus. 46 · April 30, 2021 Larochelle and Larivière Geographic distribution (Fig. 125). South Island: BR–Lake Rotoiti. Mount St Arnaud. KA–Williams Stream. MB–Judges Creek. Shroders Creek (Upper Wairau Valley). NN–Beebys Knob. Kaituna River (near Aorere). Nelson. Ecology. Lowlands, montane. Epigean. Dry forests (beech). Shaded ground; dry soil. Nocturnal; found under a log and a matagouri bush. Biology. Seasonality: September–October, January–February, June. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting technique. Hand-collecting. References. Larochelle and Larivière 2001: 130 (as “Anchomenus” arnaudensis; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus arnaudensis; list), 2016: 36 (list). Remarks. The holotype of Anchomenus arnaudensis could not be located in BMNH where Broun’s specimens can usually be found. However, there is no ambiguity regarding the identity of this taxon. Furthermore, a nontype female specimen from the Broun Collection (BMNH) was examined. This specimen is from Lake Rotoiti (BR) at St Arnaud next to the St Arnaud Range and its morphology corresponds to Broun’s original description of Anchomenus arnaudensis. In addition to diagnostic characters of the male genitalia, Ctenognathus arnaudensis has the following distinguishing features: femora and tibiae dark rufous; pronotum subrectangular, base subtruncate; elytra slightly convex, microsculpture moderately transverse, striae impunctate. Ctenognathus arnaudensis occurs in dry beech forests from northern areas of the South Island (NN, BR, MB, KA). Ctenognathus marieclaudiae Larochelle, new species Fig. 67, 110, 137 Ctenognathus marieclaudiae Larochelle, new species. Holotype: male (NZAC) labeled “NEW ZEALAND OL (typed) Makarora 3 Feb 1984 JC Watt (hand-written) / on mossy trunks at night (hand-written) / HOLOTYPE [male symbol] Ctenognathus marieclaudiae Larochelle, 2021 (red label; typed).” Paratypes: one female (NZAC) and one male (AMNZ) from the same locality as the holotype, bearing blue paratype labels. Description. Body length 12.6–13.3 mm. Head, pronotum, elytra, and abdomen black; antennae and palpi dark rufous; legs black. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric (almost granulate) on elytra. Iridescence absent. Head, pronotum, and elytra very shiny. Forebody very wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, moderately emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction absent dorsolaterally. Mentum: medial tooth slightly emarginate apically; two setae. Submentum with six setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, unwrinkled, wide, moderately cordate, widest before middle; apex subtruncate; anterolateral angles poorly developed, narrowly rounded; anterior bead complete; sides moderately rounded, slightly sinuate posteriorly; lateral beads complete; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles rectangular; laterobasal foveae moderately deep and wide, prolonged forward; posterior bead incomplete, obsolete medially; base subtruncate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Slightly convex, sloping toward apex, oblong, widest behind middle. Shoulders moderately rounded. Sides slightly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae deep, impunctate. Intervals depressed; interval 3 with three or four moderately deep, large (foveate) setiferous punctures. Umbilicate series with 17–20 setiferous punctures. Subapical sinuations moderate. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with four long apical ambulatory setae; female with ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 110): moderately arcuate, slender; base strongly convex dorsally, with basal lobe moderately wide; middle moderately convex, then rather straight toward apex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and long; apex slender, strongly convex dorsally, almost straight ventrally, Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 47 with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 63 specimens (AMNZ, BMNH, MONZ, NZAC). Geographic distribution (Fig. 137). South Island: BR, OL, WD. Ecology. Lowland, montane. Epigean, arboreal. Wet forests (beech, podocarp, broadleaf). Shaded ground; wet soil. Nocturnal; hides during the day under logs, fallen branches, and stones; active at night on mossy tree trunks and logs. Biology. Seasonality: November–April. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Pitfall trapping; turning logs, fallen branches, and stones; examining mossy tree trunks and logs at night. Remarks. The species is named after Marie-Claude Larivière, the love of my life. Ctenognathus marieclaudiae Larochelle is morphologically close to C. edwardsii. In addition to diagnostic characters of the male genitalia, C. marieclaudiae has the following distinguishing features: pronotum unwrinkled and moderately cordate, laterobasal foveae moderately wide, not meeting toward pronotal midline; elytra very shiny, slightly convex, interval 3 with three or four large (foveate) setiferous punctures. C. marieclaudiae is found in western areas of the South Island (BR, WD, OL), while C. edwardsii occurs in the southern South Island (MC to FD–SL) and Stewart Island. Ctenognathus edwardsii (Bates, 1874) Fig. 68, 111, 130 Platynus Edwardsii Bates, 1874: 239 (redescribed in 1875: 302). Holotype: “New Zealand” (could not be located either in MNHN or in CMNZ). Anchomenus (Platynus) edwardsii: Broun 1880: 26. Anchomenus macrocoelis Broun, 1908: 346. Lectotype (here designated): male (BMNH) labeled “Type (circular red-bordered label; typed) / 2664. [male symbol] (hand-written) / [MK] Hermitage. Mt. Cook. (hand-written) / New Zealand. Broun. Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / Anchomenus macrocoelis. (hand-written) / LECTOTYPE [male symbol] Anchomenus macrocoelis Broun, 1908 designated by Larochelle & Larivière, 2021 (red label; typed).” New synonym Anchomenus edwardsii: Hudson 1923: 356. Agonum (Platynus) edwardsii: Csiki 1931: 853. Agonum (Anchomenus) macrocoele: Csiki 1931: 865. Anchomenus edwardsii: Hudson 1934: 174. Anchomenus macrocaelis [sic]: Hudson 1934: 174. Ctenognathus edwardsii: Larochelle and Larivière 2007: 81, 111. Ctenognathus macrocoelis: Larochelle and Larivière 2007: 81, 111. Description. Body length 10.4–12.6 mm. Head, pronotum, elytra, and abdomen black; antennae and palpi dark rufous; legs piceous black (tarsi with reddish tinge). Microsculpture isodiametric on head, pronotum, and elytra (elytra almost granulate). Iridescence absent. Head and pronotum very shiny, elytra dull. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, moderately emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction absent dorsolaterally. Mentum: medial tooth slightly or moderately emarginate apically; two setae. Submentum with six setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc, deeply wrinkled across base, wide, subquadrate, widest before middle; apex strongly emarginate; anterolateral angles well developed, broadly rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral beads complete; lateral depressions narrow anteriorly, widened posteriorly; two setiferous punctures (anterior and posterior) on each side; posterolateral angles rectangular; laterobasal foveae moderately deep, very wide, meeting toward pronotal midline, prolonged forward; posterior bead obsolete; base subtruncate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, 48 · April 30, 2021 Larochelle and Larivière tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Subdepressed, not sloping down toward apex, oblong, widest behind middle. Shoulders moderately rounded. Sides moderately rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae shallow, impunctate. Intervals subdepressed; interval 3 with three shallow, minute setiferous punctures. Umbilicate series with 17–19 setiferous punctures. Subapical sinuations strong. Apices rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with four long apical ambulatory setae; female with eight or ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 111): strongly arcuate; base slightly convex dorsally, with marginal blade instead of basal lobe; middle slightly convex, almost straight in basal half and strongly convex in apical half dorsally, strongly concave ventrally, with dorsal membranous area very wide in apical half and moderately long; apex slender, slightly convex dorsally, slightly concave ventrally, with extreme tip narrow and very long. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 192 specimens (AMNZ, BMNH, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 130). South Island: CO, FD, MC, MK (mostly), OL, SC, SL. Stewart Island. Ecology. Lowland, montane, subalpine. Epigean. Dry forests (beech). Shaded ground; dry soil. Nocturnal; hides during the day under logs and stones. Gregarious. Biology. Seasonality: October–February, April–May. Tenerals: November, January–March, May. Predacious (based on mouthpart morphology). Occasionally infested with mites. Dispersal power. Subapterous (incapable of flight). Moderate runner. Occasional climber (on trees). Collecting techniques. Turning logs and stones; pitfall trapping. References. Larochelle and Larivière 2001: 130–131 (as “Anchomenus” edwardsii, “A.” macrocoelis; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus edwardsii, C. macrocoelis; list), 2016: 36 (as Ctenognathus edwardsii, C. macrocoelis; list). Remarks. The holotype of Platynus edwardsii could not be located either in MNHN or in CMNZ where Bates’ specimens can usually be found. However, there is no ambiguity regarding the identity of this taxon; the original description can only apply to the species as currently understood. In addition to diagnostic characters of the male genitalia, Ctenognathus edwardsii has the following distinguishing features: pronotum wrinkled on disc and across base, subquadrate, laterobasal foveae of pronotum very wide, meeting toward pronotal midline; elytra dull, subdepressed, interval 3 with three shallow, minute setiferous punctures. Broun described Anchomenus macrocoelis from two specimens, one of which (a male) was located (BMNH) and is here designated as a lectotype in order to preserve stability of nomenclature in the future. The morphology of this lectotype corresponds to Bates’ original description of Platynus edwardsii. The two taxa are therefore conspecific. Ctenognathus kaikoura Larochelle and Larivière, new species Fig. 69, 112, 136 Ctenognathus kaikoura Larochelle and Larivière, new species. Holotype: male (NZAC) labeled “[NEW ZEALAND KA] Puhipuhi Val. [=Puhi Puhi Valley] Kaikoura Marlborough (typed) / 31 Aug 70. G.W. Ramsay Under stones. (hand-written) / HOLOTYPE [male symbol] Ctenognathus kaikoura Larochelle & Larivière, 2021 (red label; typed).” Paratypes: one female (NZAC) from Hundalee (KA) and one female (LUNZ) from Oaro (KA), bearing blue paratype labels. Description. Body length 9.8–10.5 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen smoky brown, yellowish lateroapically; antennae and palpi rufotestaceous; legs reddish. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Iridescence absent. Very shiny. Forebody narrow in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate or subtriangular apically; two setae. Submentum with six setae. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 49 Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc and across base, narrow, moderately cordate, widest before middle; apex slightly emarginate; anterolateral angles well developed, rounded; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral beads complete; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles acute, strongly projected laterally; laterobasal foveae very deep, moderately wide, prolonged forward; posterior bead complete; base subtruncate. Legs. Very long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 poorly developed, not carinate dorsally, shallowly bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Subdepressed, sloping down toward apex, subovate, widest about middle. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae shallow, impunctate. Intervals subdepressed; interval 3 with three moderately deep, minute setiferous punctures. Umbilicate series with 16–19 setiferous punctures. Subapical sinuations feeble. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with eight long apical ambulatory setae. Aedeagus. Lateral view (Fig. 112): strongly arcuate; base moderately convex dorsally, with basal lobe narrow; middle strongly convex with a slight concavity before apex dorsally, moderately concave ventrally, with dorsal membranous area strongly widened in apical half and moderately long; apex subtriangular, slightly convex dorsally, slightly concave ventrally, with extreme tip subtriangular and very long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 7 specimens (AMNZ, BMNH, LUNZ, NZAC). Geographic distribution (Fig. 136). South Island: KA–Hundalee. Oaro. Puhi Puhi Valley. MB–Saxton Pass. Wairau River Bridge (above Judges Creek). Ecology. Lowland, montane. Dry forests (beech). Shaded ground; dry soil. Nocturnal; hides during the day under stones and in leaf litter. Biology. Seasonality: October, December, February, August. Tenerals: March. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Swift runner. Collecting techniques. Lifting stones; sifting leaf litter. Remarks. The species is named the Kaikoura region (KA) which includes the type locality Puhi Puhi Valley, and is applied as a noun in apposition. In addition to diagnostic characters of the male genitalia, Ctenognathus kaikoura has mesoand metatarsomeres 1–4 without dorsal carinae, characters that separate it from other South Island Ctenognathus species. Ctenognathus kaikoura is restricted to the dry beech forests of northeastern areas of the South Island (MB, KA). Ctenognathus sandageri (Broun, 1882) Fig. 70, 113, 143 Anchomenus sandageri Broun, 1882: 218 (redescribed in 1883: 218 and 1886: 748). Holotype: male (BMNH) labeled “[male symbol] (hand-written) / Type (circular red-bordered label; typed) / 46. (hand-written) / New Zealand Broun. Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / [WN] Wellington (typed) / Anchomenus sandageri. (hand-written).” Agonum (Platynus) sandageri: Csiki 1931: 853. Anchomenus sandageri: Hudson 1934: 174. Ctenognathus sandageri: Larochelle and Larivière 2007: 81, 111. Description. Body length 10.2–11.1 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen piceous black, yellowish lateroapically; antennae and palpi rufotestaceous; femora and tibiae pale yellow; tarsi rufotestaceous. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Iridescence absent. Very shiny. Forebody narrow in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each 50 · April 30, 2021 Larochelle and Larivière eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate or subrounded apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum moderately convex, impunctate, obsoletely wrinkled on disc, narrow, strongly cordate, widest about middle; apex strongly emarginate; anterolateral angles poorly developed, rounded; anterior bead complete; sides strongly rounded, strongly sinuate posteriorly; lateral beads complete; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles acute, slightly projected laterally; laterobasal foveae moderately deep and wide, prolonged forward; posterior bead incomplete, obsolete medially; base emarginate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Very convex, sloping down toward apex, subovate, widest about middle. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae deep, impunctate. Intervals subdepressed; interval 3 with three moderately deep, small setiferous punctures. Umbilicate series with 15–16 setiferous punctures. Subapical sinuations moderate. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 113): moderately arcuate, strongly widened in apical half; base moderately convex dorsally, with basal lobe narrow; middle moderately convex dorsally, strongly concave ventrally, with dorsal membranous area strongly widened in apical half and moderately long; apex subtriangular, moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 201 specimens (AMNZ, BMNH, CMNZ, JNNZ, MONZ, NZAC). Geographic distribution (Fig. 143). North Island: WA, WN. Ecology. Lowland, montane. Epigean. Wet forests (broadleaf, beech, podocarp). Associated with streams. Shaded ground; wet soil. Nocturnal; hides during the day under stones and logs. Gregarious. Biology. Seasonality: October–March. Tenerals: November, March, May. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Pitfall trapping; turning stones and logs. References. Larochelle and Larivière 2001: 131 (as “Anchomenus” sandageri; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus sandageri), 2016: 37 (list). Ctenognathus helmsi (Sharp, 1881) Fig. 71, 114, 133 Anchomenus helmsi Sharp, 1881: 47. Lectotype (here designated): female (BMNH) labeled “Anchomenus helmsi [WD] Greymouth N. Zd. (hand-written) / Greymouth New Zealand. Helms. (white label with red horizontal line; typed) Sharp Coll. 1905–313. / LECTOTYPE [female symbol] Anchomenus helmsi Sharp, 1881 designated by Larochelle & Larivière, 2021 (red label; typed).” Agonum (Platynus) helmsi: Csiki 1931: 853. Anchomenus helmsi: Hudson 1934: 174. Agonum helmsi: Johns 1977: 316. Anchomenus helmsi: Townsend 1997: 15. Ctenognathus helmsi: Larochelle and Larivière 2007: 81, 111. Description. Body length 9.5–11.6 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen piceous black, yellowish lateroapically; antennae and palpi rufotestaceous; femora dark yellow; tibiae and tarsi reddish. Microsculpture isodiametric on head, moderately transverse on pronotum, and isodiametric on elytra. Iridescence absent. Very shiny. Forebody narrow in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 51 not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate or slightly emarginate apically; two setae. Submentum with four setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum moderately convex, impunctate, obsoletely wrinkled throughout, narrow, strongly cordate, widest about middle; apex strongly emarginate; anterolateral angles well developed, rounded; anterior bead complete; sides strongly rounded, strongly sinuate posteriorly; lateral beads complete; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles acute, slightly projected laterally; laterobasal foveae very deep, moderately wide, prolonged forward; posterior bead complete; base subtruncate. Apex of prosternum glabrous or pubescent (glabrous in all other species). Legs. Very long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 setose ventrally (with very short setae). Elytra. Moderately convex, sloping down toward apex, subovate, widest about middle. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae deep, impunctate. Intervals subdepressed; interval 3 with three moderately deep, small setiferous punctures. Umbilicate series with 16–18 setiferous punctures. Subapical sinuations strong. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 114): moderately arcuate; base strongly convex dorsally, with basal lobe narrow; middle moderately convex dorsally, strongly concave ventrally, with dorsal membranous area strongly widened in apical half and very long; apex slender, slightly convex dorsally, slightly concave ventrally, with extreme tip subtriangular and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 322 specimens (AMNZ, BMNH, CMNH, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 133). South Island: BR, NN, SD, WD. Ecology. Lowland, montane. Epigean. Wet forests (beech, broadleaf, podocarp). Associated with streams and mud flats. Shaded ground; wet soil. Nocturnal; hides during the day under logs and stones. Gregarious. Biology. Seasonality: September–March, May. Tenerals: November, February–May. Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Swift runner. Collecting techniques. Turning logs and stones; pitfall trapping. References. Larochelle and Larivière 2001: 130 (as “Anchomenus” helmsi; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus helmsi; list), 2016: 37 (list). Remarks. Sharp described Anchomenus helmsi from two specimens, one of which (a female) was located (BMNH) and is here designated as lectotype. This type designation is made to preserve stability of nomenclature in the future. Ctenognathus xanthomelus (Broun, 1908) Fig. 72, 115, 148 Anchomenus xanthomelus Broun, 1908: 346. Holotype: male (BMNH) labeled “Type (circular red-bordered label; typed) / 2665. [male symbol] (hand-written) / New Zeal. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; typed) / [RI/WN] Manawatu Gorge. (hand-written) / Anchomenus xanthomelus (hand-written).” Anchomenus xanthomelas [sic]: Hudson 1923: 356. Agonum (Anchomenus) xanthomelus: Csiki 1931: 865. Anchomenus xanthomelas [sic]: Hudson 1934: 174. Ctenognathus xanthomelus: Larochelle and Larivière 2007: 81, 111. Description. Body length 10.0–11.4 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen piceous black, yellow lateroapically; antennae and palpi rufotestaceous; femora pale yellow; tibiae and tarsi reddish. Microsculpture isodiametric on head, very transverse (with microlines) on pronotum, and moderately transverse on elytra. Iridescence present on pronotum. Very shiny. Forebody narrow in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. 52 · April 30, 2021 Larochelle and Larivière Labrum strongly transverse, slightly emarginate anteriorly. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth slightly emarginate apically; two setae. Submentum with six setae. Palpi with terminal segment truncate apically. Thorax. Pronotum moderately convex, impunctate, deeply wrinkled across base, narrow, strongly cordate, widest about middle; apex slightly emarginate; anterolateral angles poorly developed, rounded; anterior bead complete; sides strongly rounded, strongly sinuate posteriorly; lateral beads absent; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles rectangular; laterobasal foveae very deep, moderately wide, prolonged forward; posterior bead complete; base subtruncate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Very convex, sloping down toward apex, ovate, widest about middle. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae deep, finely punctate. Intervals slightly convex; interval 3 with three moderately deep, small setiferous punctures. Umbilicate series with 16–18 setiferous punctures. Subapical sinuations feeble. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 115): moderately arcuate, strongly widened in apical half; base moderately convex dorsally, with basal lobe narrow; middle moderately concave dorsally, strongly concave ventrally, with dorsal membranous area moderately wide and moderately long; apex subtriangular, strongly convex dorsally, slightly concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 127 specimens (AMNZ, BMNH, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 148). North Island: BP, RI, TK, TO, WN, WO. Ecology. Lowland, montane. Epigean. Wet forests (broadleaf, podocarp). Associated with streams, slopes, and mud flats. Shaded ground; wet soil. Nocturnal; hides during the day under logs and fallen branches, also in heaps of dead leaves and mud. Gregarious. Biology. Seasonality: October–January, March–April, June. Tenerals: November–December, February, June. Predacious (based on mouthpart morphology). Occasionally infested with mites. Dispersal power. Subapterous (incapable of flight). Moderate runner. Collecting techniques. Pitfall trapping; turning logs and fallen branches; examining heaps of dead leaves and mud; pan trapping (yellow pan traps). References. Larochelle and Larivière 2001: 132 (as “Anchomenus” xanthomelas; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus xanthomelus; list), 2016: 38 (list). Ctenognathus intermedius (Broun, 1908) Fig. 73, 116, 135 Anchomenus intermedius Broun, 1908: 347. Lectotype (here designated): male (BMNH) labeled “Type (circular red-bordered label; typed) / 2666. [male symbol] (hand-written) / [WI/WN] Manawatu Flats (hand-written) / Anchomenus intermedius. (hand-written) / New Zealand. Broun. Coll. Brit. Mus. 1922–482. (white label with red horizontal line; typed) / LECTOTYPE [male symbol] Anchomenus intermedius Broun, 1908 designated by Larochelle & Larivière, 2021 (red label; typed).” Agonum (Anchomenus) intermedium: Csiki 1931: 865. Anchomenus intermedius: Hudson 1934: 174. Ctenognathus intermedius: Larochelle and Larivière 2007: 81, 111. Description. Body length 10.5–12.0 mm. Head, pronotum, and elytra mostly piceous black; lateral margins of pronotum and elytra reddish; abdomen piceous black, yellowish lateroapically; antennae and palpi rufotestaceous; femora and tibiae pale yellow; tarsi reddish. Microsculpture isodiametric on head, very transverse (with Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 53 microlines) on pronotum, and isodiametric on elytra. Iridescence present on pronotum. Very shiny. Forebody narrow in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, slightly emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate apically; two setae. Submentum with six setae. Palpi with terminal segment truncate apically. Thorax. Pronotum moderately convex, impunctate, obsoletely wrinkled on disc and across base, narrow, moderately cordate, widest before middle; apex slightly emarginate; anterolateral angles poorly developed, obtuse; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral beads absent; lateral depressions wide; two setiferous punctures (anterior and posterior) on each side; posterolateral angles rectangular; laterobasal foveae very deep, moderately wide, prolonged forward; posterior bead complete; base truncate. Legs. Very long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Slightly convex, not sloping down toward apex, subovate, widest about middle. Shoulders strongly rounded. Sides strongly rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae deep, finely punctate. Intervals subdepressed; interval 3 with three moderately deep, small setiferous punctures. Umbilicate series with 17–19 setiferous punctures. Subapical sinuations feeble. Apices obtuse. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with two long apical ambulatory setae; female with four long apical ambulatory setae. Aedeagus. Lateral view (Fig. 116): moderately arcuate, slender; base moderately convex dorsally, with basal lobe narrow; middle subparallel, moderately convex dorsally, moderately concave ventrally, with dorsal membranous area moderately wide and moderately long; apex moderately convex dorsally, slightly concave ventrally, with extreme tip narrow and moderately long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 54 specimens (BMNH, JNNZ, MONZ, NZAC). Geographic distribution (Fig. 135). North Island: RI, TO, WI, WN. Ecology. Lowland, montane. Epigean. Wet forests (beech, broadleaf). Associated with streams. Shaded ground; wet soil. Nocturnal; hides during the day under logs, fallen branches, and stones. Gregarious. Biology. Seasonality: September–October, December–February. Mating observed in January (WI). Tenerals: February–April. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Swift runner. Collecting techniques. Turning logs, fallen branches, and stones. References. Larochelle and Larivière 2001: 130 (as “Anchomenus” intermedius; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus intermedius), 2016: 37 (list). Remarks. Broun described Anchomenus intermedius from two specimens, one of which (a male) could be located (BMNH) and is here designated as lectotype. This type designation is made to preserve stability of nomenclature in the future. Ctenognathus colensonis (White, 1846) Fig. 75, 117, 127 Anchomenus colensonis White, 1846: 3. Lectotype (here designated): female (BMNH) labeled “[female symbol] (handwritten) / [?]5 [first number unreadable] (hand-written) / [NC/MC/SC] Canterbury (typed) / New Zeal. Broun Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / Platynus colensonis (hand-written).” LECTOTYPE [female symbol] Anchomenus colensonis White, 1846 designated by Larochelle & Larivière, 2021 (red label; typed).” Platynus colensonis: Bates 1874: 239. Anchomenus (Platynus) colensonis: Broun 1880: 26. Anchomenus colensonis: Sharp 1884: 298. Anchomenus integratus Broun, 1908: 348. Holotype: female (BMNH) labeled “Type (circular red-bordered label; typed) / 2667 (hand-written) / New Zeal. Broun. Coll. Brit. Mus. 1922–482 (white label with red horizontal line; typed) / [MC] Broken River (hand-written) / Anchomenus integratus. (hand-written).” New synonym 54 · April 30, 2021 Larochelle and Larivière Agonum (Platynus) colensoni [sic]: Csiki 1931: 865. Agonum (Anchomenus) integratum: Csiki 1931: 865. Anchomenus colensonis: Hudson 1934: 174. Agonum integratus [sic]: Johns 1980: 62. Ctenognathus colensonis: Larochelle and Larivière 2007: 81, 111. Ctenognathus integratus: Larochelle and Larivière 2007: 81, 111. Description. Body length 10.8–11.2 mm. Head, pronotum, and elytra piceous black; lateral margins of elytra reddish; abdomen piceous black; antennae and palpi rufotestaceous; femora and tibiae pale yellow; tarsi reddish. Microsculpture isodiametric on head, pronotum, and elytra. Iridescence absent. Very shiny. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum strongly transverse, deeply emarginate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction shallow dorsolaterally. Mentum: medial tooth subtruncate apically; two setae. Submentum with ten setae. Palpi with terminal segment truncate apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc, wide, slightly cordate, widest before middle; apex slightly emarginate; anterolateral angles well developed, broadly rounded; anterior bead complete; sides moderately rounded, slightly or moderately sinuate posteriorly; lateral beads complete; lateral depressions narrow anteriorly, widened posteriorly; two setiferous punctures (anterior and posterior) on each side; posterolateral angles slightly obtuse, almost rectangular; laterobasal foveae shallow, moderately wide, prolonged forward; posterior bead obsolete; base subtruncate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Slightly convex, not sloping down toward apex, oblong, widest about middle. Shoulders moderately rounded. Sides moderately rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae shallow, finely punctate. Intervals depressed; interval 3 with three shallow, minute setiferous punctures. Umbilicate series with 15–22 setiferous punctures. Subapical sinuations feeble. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with four long apical ambulatory setae; female with eight or ten long apical ambulatory setae. Aedeagus. Lateral view (Fig. 117): strongly arcuate; base strongly convex dorsally, with basal lobe moderately wide; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area very wide in apical half and moderately long; apex moderately convex dorsally, slightly concave ventrally, with extreme tip moderately wide and long. Dorsal view: very wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 363 specimens (AMNZ, BMNH, CMNH, JNNZ, LUNZ, NZAC). Geographic distribution (Fig. 127). South Island: BR, MC, MK, NC, NN, SC. Ecology. Lowland, montane, subalpine. Epigean, arboreal. Dry forests (beech). Shaded ground; dry soil. Nocturnal; hides during the day under logs and fallen branches. Gregarious. Biology. Seasonality: September–May, July. Tenerals: November–December, February–March, May. Predacious (based on mouthpart morphology). Dispersal power. Subapterous (incapable of flight). Moderate runner. Regular climber (on tree trunks). Collecting techniques. Pitfall trapping; turning logs and fallen branches; examining tree trunks at night. References. Larochelle and Larivière 2001: 130 (as “Anchomenus” colensonis, “A.” integratus; catalogue; biology, dispersal power, ecology, geographic distribution, references), 2007: 81, 111 (as Ctenognathus colensonis, C. integratus; list), 2016: 36–37 (as C. colensonis, C. integratus; list). Remarks. White described Anchomenus colensonis from two specimens, one of which (a female) could be located (BMNH) and is here designated as lectotype in order to preserve stability of nomenclature in the future. Examination of the type of Anchomenus integratus revealed it to be conspecific with Ctenognathus colensonis. Ctenognathus otagoensis (Bates, 1878) Fig. 74, 118, 140 Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 55 Anchomenus (Platynus) otagoensis Bates, 1878: 27. Syntypes: “[male symbol] [female symbol]” from [CO/DN/OL] “Otago” (could not be located either in MNHN or in CMNZ). Anchomenus otagoensis: Broun 1880: 25. Agonum (Platynus) otagoense: Csiki 1931: 853. Platynus otagoensis: Liebherr and Zimmerman 1998: 147. Ctenognathus otagoensis: Liebherr 2005: 283. Description. Body length 10.3–12.7 mm. Head, pronotum, and elytra piceous black; abdomen rufopiceous; antenna and palpi rufotestaceous; femora and tibiae pale yellow; tarsi reddish. Microsculpture isodiametric on head, pronotum, and elytra (elytra almost granulate). Iridescence absent. Head and pronotum very shiny, elytra dull. Forebody moderately wide in comparison to elytra. Head. Moderately wide. Mandibles moderately long and curved anteriorly. Labrum moderately transverse, subtruncate anteriorly. Eyes strongly convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Neck constriction absent dorsolaterally. Mentum: medial tooth slightly or moderately emarginate apically; two setae. Submentum with six setae. Palpi with terminal segment obtuse apically. Thorax. Pronotum slightly convex, impunctate, obsoletely wrinkled on disc, wide, moderately cordate, widest before middle; apex slightly emarginate; anterolateral angles poorly developed, obtuse; anterior bead incomplete, obsolete medially; sides moderately rounded, slightly sinuate posteriorly; lateral beads complete; lateral depressions narrow anteriorly, widened posteriorly; two setiferous punctures (anterior and posterior) on each side; posterolateral angles rectangular; laterobasal foveae moderately deep and wide, prolonged forward; posterior bead complete; base truncate. Legs. Moderately long. Metafemora with two posteroventral setae. Mesoand metatarsomeres 1–4 well developed, tricarinate dorsally, deeply bisulcate laterally. Metatarsomeres 4 moderately bilobed, symmetrical apically. Metatarsomeres 5 glabrous ventrally. Elytra. Subdepressed, not sloping down toward apex, oblong, widest about middle. Shoulders moderately rounded. Sides moderately rounded. Scutellar setiferous pore visible. Scutellar striole moderately long, impunctate. Striae shallow, finely punctate. Intervals depressed; interval 3 with three shallow, minute setiferous punctures. Umbilicate series with 17–20 setiferous punctures. Subapical sinuations strong. Apices obtusely rounded. Abdomen. Sterna IV–VI: both sexes with two long apical ambulatory setae. Sternum VII (last visible sternum): male with four long apical ambulatory setae; female with eight long apical ambulatory setae. Aedeagus. Lateral view (Fig. 118): strongly arcuate, strongly widened medially; base strongly convex dorsally, with marginal blade instead of basal lobe; middle strongly convex dorsally, slightly convex ventrally, with dorsal membranous area strongly widened in apical half and moderately long; apex strongly convex dorsally, moderately concave ventrally, with extreme tip narrow and very long. Dorsal view: moderately wide, asymmetrical (ostium of membranous area deflected to the left); basal bulb moderately distant from membranous area. Material examined. 615 specimens (AMNZ, JNNZ, LUNZ, MONZ, NZAC). Geographic distribution (Fig. 140). South Island: CO, DN, FD, OL, SL. Ecology. Lowland, montane, subalpine. Epigean. Wet forests (beech, broadleaf), tree plantations (pine), shrublands, and scrublands. Associated with streams. Shaded ground; wet soil. Nocturnal; hides during the day under logs, fallen trees, and stones. Gregarious. Biology. Seasonality: September–April, June, August. Tenerals: March. Predacious (based on mouthpart morphology). Occasionally infested with fungi (Laboulbeniales). Dispersal power. Subapterous (incapable of flight). Moderate runner. Occasional climber (on trees). Collecting techniques. Pitfall trapping; turning logs, fallen trees, and stones. References. Larochelle and Larivière 2001: 131 (as “Anchomenus” otagoensis; catalogue; biology, dispersal power, ecology, geographic distribution, references); Liebherr 2005: 283 (as Ctenognathus otagoensis); Larochelle and Larivière 2007: 111 (list), 2016: 37 (list). Remarks. The syntypes of Anchomenus otagoensis could not be located either in MNHN or in CMNZ where Bates’ specimens can usually be found. However, there is no ambiguity regarding the identity of this taxon; the original description can only apply to the species as currently understood. 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Akatarawa Range WN −40.9500 175.1000 Alex Knob WD −43.4277 170.1513 Aorere NN −40.7221 172.6007 Auckland AK −36.8525 174.7622 Aupouri Peninsula ND −34.7000 173.0000 Banks Peninsula MC −43.8825 173.0089 Bay of Islands ND −35.2166 174.2000 Beebys Knob NN −41.7329 172.9315 Big South Cape Island SI −47.2377 167.4075 Broken River MC −43.2006 171.7881 Callary Gorge WD −43.3931 170.1874 Canterbury NC/MC/SC −43.0000 172.0000 Cape Maria Van Dimmen ND −34.4758 172.6433 Castaway Camp, Great Island TH −34.1534 172.1413 Catlins River Track SL −46.4534 169.4685 Catlins State Forest Park SL −46.4800 169.3300 Chatham Islands CH −44.0000 176.5000 Christchurch MC −43.5160 172.5242 Clevedon AK −36.9923 175.0375 East Cape GB −37.7877 178.5492 Fiordland National Park FD −45.3800 167.2800 Fox Glacier WD −43.4660 170.0199 Frank Stream, Catlins State Forest Park SL −46.4426 169.4623 Franz Josef WD −43.3887 170.1824 Great Island TH −34.1666 172.1333 Greymouth WD −42.4610 171.2046 Hakapoua FD −46.1666 166.9500 64 · April 30, 2021 Larochelle and Larivière Hermitage, Mount Cook MK −43.7333 170.0833 Homer Saddle FD −44.7593 167.9798 Huiarau Summit GB −38.6166 177.0166 Huka Stream ND −34.4286 172.9343 Hundalee KA −42.5988 173.4191 Hunua Ranges AK −37.0750 175.1800 Invercargill SL −46.4166 168.3666 Judges Creek MB −42.0557 172.9635 Kaituna River, near Aorere NN −40.7221 172.6007 Kaituna Stream/Valley, Banks Peninsula MC −43.7292 172.6985 Karangarua River WD −43.5765 169.8092 Lake Rotoiti BR −41.8333 172.8333 Lake Wilkie SL −46.5801 169.4391 Lake Wombat WD −43.4032 170.1676 Macdonalds Creek WD −43.2953 170.2283 Makarora OL −44.2332 169.2328 Manawatu Gorge RI/WN −40.3000 175.7666 Manawatu River Flats WI/WN −40.4700 175.2000 Mangamuka Gorge Walkway ND −35.1917 173.4550 Matapia Island ND −34.6060 172.7984 Mataraua Forest ND −35.5728 173.6506 Maungapiko Hill ND −34.4213 172.8622 Middle Road, Banks Peninsula MC −43.7166 172.6833 Mokau Road to Kiwi Road Track TK −38.9114 174.6069 Mokau Road to White Cliffs Track TK −38.8915 174.5874 Motuopao Island ND −34.4702 172.6377 Mount Gray, Turret Range FD −45.5526 167.2480 Mount Manaia ND −35.8181 174.5171 Mount Maungatua DN −45.8833 170.1166 Mount Messenger TK −38.9000 174.9333 Mount Quoin WN −41.0000 175.2333 Murimotu Island ND −34.4144 173.0500 Nelson NN −41.2734 173.2823 Ngaroku Stream ND −34.4133 173.0319 Oaro KA −42.5164 173.5044 Okarito WD −43.2232 170.1616 Onepu Hut Track, Waimana Valley BP −38.3264 177.1195 Otago CO/DN/OL −45.5000 170.0000 Parua ND −35.7669 174.4716 Picton SD −41.2922 174.0011 Puhi Puhi Valley KA −42.2826 173.7398 Remuera, Auckland AK −36.8763 174.7994 Reynolds, Banks Peninsula MC −43.7833 172.8167 Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 65 Saxton Pass MB −42.0696 173.2196 Shroders Creek MB −41.9879 172.8812 Slopedown Range SL −46.3500 169.0800 Spirits Bay ND −34.4588 172.8056 St Arnaud BR −41.8038 172.8470 Stewart Island SI −46.9962 167.8842 Stratford TK −39.3408 174.2844 Takahe Valley FD −45.2891 167.6574 Tasman Valley, Great Island TH −34.1608 172.1372 Taumaka Island/Forest WD −43.8601 168.8832 Tawanui SL −46.4622 169.5255 Te Kuiti WO −38.3333 175.1666 Te Paki ND −34.5039 172.7991 Te Waiiti Stream, Waimana Valley BP −38.3084 177.1355 Unuwhao ND −34.4321 172.8882 Urewera National Park GB −38.7500 177.1500 Waimana Valley BP −38.2424 177.1085 Waipoua River, Waipoua Forest ND −35.6524 173.5595 Waipuna Cave, Te Kuiti WO −38.2500 175.0833 Wairau River Bridge, Judges Creek MB −42.0665 172.9308 Wairoa GB −39.0417 177.4233 Wairoa, Hunua Ranges AK −37.0746 175.1361 Waitakere Railway-station AK −36.8507 174.5434 Wallis Stream, Catlins State Forest Park SL −46.4286 169.4507 Wekaweka Road end, Mataraua Forest ND −35.5607 173.6058 Wellington WN −41.2783 174.7771 West Plains SL −46.3680 168.3149 Whangarei Heads ND −35.8133 174.5055 Williams Stream KA −42.1132 173.7453 66 · April 30, 2021 Larochelle and Larivière Figures 1–13. Microsculpture: 1) granulate; 2) isodiametric; 3) moderately transverse; 4) very transverse. Neck constriction, dorsolateral view: 5) deep; 6) shallow; 7) absent. Eye and buccal fissure, lateral view: 8) touching; 9) separated. Terminal segment of palpi: 10) obtuse apically; 11) truncate. Paraglossae: 12) pubescent; 13) glabrous. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 67 Figures 14–26. Mentum: 14) medial tooth bifid; 15) emarginate; 16) subtruncate; 17) subtriangular; 18) acute; 19) circular foveae large; 20) small; 21) with numerous short setae; 22) without short setae. Metacoxae: 23) with 3–4 posteroventral setae; 24) with 2 setae. Cleaning organ of protibia: 25) with 2 clip setae; 26) with 3 clip setae. 68 · April 30, 2021 Larochelle and Larivière Figures 27–36. Mesoor metatarsomere, dorsal view: 27) not carinate; 28) tricarinate. Metatarsomere 4: 29) moderately bilobed and asymmetrical apically; 30) moderately bilobed and symmetrical; 31) strongly bilobed and asymmetrical. Abdominal sterna V–VI: 32) with 4 ambulatory setae; 33) with 2 ambulatory setae; 34) V–VII with numerous short setae; 35) VII, male with 2 ambulatory setae, female with 4 setae; 36) VII, male with 4 ambulatory setae, female with 8 setae. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 69 Figures 37–40. Dorsal habitus. 37) Platynus macropterus. 38) Notagonum chathamense. 39) N. lawsoni. 40) N. feredayi. Scale line = 1 mm. 70 · April 30, 2021 Larochelle and Larivière Figures 41–44. Dorsal habitus. 41) Notagonum submetallicum. 42) Prosphodrus mangamuka new species. 43) P. waltoni. 44) P. sirvidi new species. Scale line = 1 mm. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 71 Figures 45–48. Dorsal habitus. 45) Prosphodrus waimana new species. 46) P. occultus. 47) Maoriplatynus marrisi new species. 48) Kiwiplatynus bidens. Scale line = 1 mm. 78 · April 30, 2021 Larochelle and Larivière Figures 73–76. Dorsal habitus. 73) Ctenognathus intermedius. 74) C. otagoensis. 75) C. colensonis. 76) C. earlyi new species. Scale line = 1 mm. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 79 Figures 77–79. Dorsal habitus. 77) Ctenognathus davidsoni new species. 78) C. oreobius. 79) C. hoarei new species. Scale line = 1 mm. 80 · April 30, 2021 Larochelle and Larivière Figures 80–86. Aedeagus, lateral. 80) Platynus macropterus. 81) Notagonum chathamense. 82) N. lawsoni. 83) N. feredayi. 84) N. submetallicum. 85) Prosphodrus mangamuka new species. 86) P. waltoni. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 81 Figures 87–93. Aedeagus, lateral. 87) Prosphodrus sirvidi new species. 88) P. waimana new species. 89) P. occultus. 90) Maoriplatynus marrisi new species. 91) Kiwiplatynus bidens. 92) K. taranaki new species. 93) Kupeplatynus sulcitarsis. 82 · April 30, 2021 Larochelle and Larivière Figures 94–100. Aedeagus, lateral. 94) Kupeplatynus crenatus. 95) K. lucifugus. 96) Tuiplatynus sophronitis. 97) T. libitus. 98) Ctenognathus novaezelandiae. 99) C. elevatus. 100) C. garnerae new species. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 83 Figures 101–107. Aedeagus, lateral. 101) Ctenognathus tepaki new species. 102) C. urewera new species. 103) C. cardiophorus. 104) C. adamsi. 105) C. pictonensis. 106) C. tawanui new species. 107) C. perumalae new species. 84 · April 30, 2021 Larochelle and Larivière Figures 108–114. Aedeagus, lateral. 108) Ctenognathus takahe new species. 109) C. arnaudensis. 110) C. marieclaudiae new species. 111) C. edwardsii. 112) C. kaikoura new species. 113) C. sandageri. 114) C. helmsi. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 85 Figures 115–122. Aedeagus, lateral. 115) Ctenognathus xanthomelus. 116) C. intermedius. 117) C. colensonis. 118) C. otagoensis. 119) C. earlyi new species. 120) C. davidsoni new species. 121) C. oreobius. 122) C. hoarei new species. 86 · April 30, 2021 Larochelle and Larivière Figure 123. Map of New Zealand, outlying islands, areas and area codes. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 87 Figures 124–129. Species distribution maps. 124) Ctenognathus adamsi. 125) C. arnaudensis. 126) C. cardiophorus. 127) C. colensonis. 128) C. davidsoni new species. 129) C. earlyi new species. 94 · April 30, 2021 Larochelle and Larivière Figures 165–166. Species distribution maps. 165) Tuiplatynus libitus. 166) T. sophronitis. Synopsis of the tribe Platynini in New Zealand Insecta Mundi 0864 · 95 Taxonomic Index Names in bold indicate valid taxa. actochares Broun, Ctenognathus 34 adamsi Broun, Anchomenus 40 adamsi (Broun), Ctenognathus 40 angusticollis Fabricius, Carabus 6 arnaudensis Broun, Anchomenus 45 arnaudensis (Broun), Ctenognathus 45 batesi Broun, Anchomenus 20 bidens Chaudoir, Colpodes 20 bidens (Chaudoir), Kiwiplatynus 20 cardiophorus Chaudoir, Colpodes 38 cardiophorus (Chaudoir), Ctenognathus 38 Cerabilia Laporte de Castelnau 2 chathamense (Broun), Notagonum 8 chathamensis Broun, Anchomenus 8 cheesemani Broun, Anchomenus 24 colensonis White, Anchomenus 53 colensonis (White), Ctenognathus 53 crenatus Chaudoir, Colpodes 24 crenatus (Chaudoir), Kupeplatynus 24 Ctenognathus Fairmaire 29 davidsoni new species, Ctenognathus 57 deformipes Broun, Calathus 24 earlyi new species, Ctenognathus 56 edwardsii (Bates), Ctenognathus 47 Edwardsii Bates, Platynus 47 elevatus White, Anchomenus 34 elevatus (White), Ctenognathus 34 externum Darlington, Notagonum 7 Feredayi Bates, Anchomenus 10 feredayi (Bates), Notagonum 10 garnerae new species, Ctenognathus 35 haasti Broun, Anchomenus 6 helmsi Sharp, Anchomenus 50 helmsi (Sharp), Ctenognathus 50 hoarei new species, Ctenognathus 59 integratus Broun, Anchomenus 54 intermedius Broun, Anchomenus 52 intermedius (Broun), Ctenognathus 52 kaikoura new species, Ctenognathus 48 Kiwiplatynus new genus 19 Kupeplatynus new genus 22 Laemostenus Bonelli 2 latipennis Sharp, Ctenognathus 33 Lawsoni Bates, Anchomenus 9 lawsoni (Bates), Notagonum 9 libitus Broun, Anchomenus 28 libitus (Broun), Tuiplatynus 28 littorellus Broun, Ctenognathus 40 lucifugus Broun, Anchomenus 25 lucifugus (Broun), Kupeplatynus 25 macrocoelis Broun, Anchomenus 47 macropterus Chaudoir, Colpodes 6 macropterus (Chaudoir), Platynus 6 mangamuka new species, Prosphodrus 13 Maoriplatynus new genus 18 marginellum (Erichson), Notagonum 2 marginicolle (Macleay), Notagonum 12 marieclaudiae new species, Ctenognathus 46 marrisi new species, Maoriplatynus 18 montivagus Broun, Anchomenus 32 munroi Broun, Anchomenus 25 neozelandicus Chaudoir, Colpodes 32 Notagonum Darlington 7 Novae-Zelandiae Fairmaire, Anchomenus 32 novaezelandiae (Fairmaire), Ctenognathus 32 occultus Britton, Prosphodrus 17 oreobius Broun, Anchomenus 58 oreobius (Broun), Ctenognathus 58 otagoensis Bates, Anchomenus 55 otagoensis (Bates), Ctenognathus 55 ovicollis Motschulsky, Dicrochile 32 parabilis Broun, Anchomenus 38 perrugithorax Broun, Anchomenus 32 perumalae new species, Ctenognathus 43 pictonensis Sharp, Ctenognathus 41 Platynus Bonelli 6 politulus Broun, Anchomenus 32 Prosphodrini 13 Prosphodrus Britton 12 punctulatus Broun, Anchomenus 32 sandageri Broun, Anchomenus 49 sandageri (Broun), Ctenognathus 49 simmondsi Broun, Ctenognathus 41 96 · April 30, 2021 Larochelle and Larivière sirvidi new species, Prosphodrus 15 sophronitis Broun, Anchomenus 27 sophronitis (Broun), Tuiplatynus 27 Sphodrini 2 submetallicum (White), Notagonum 11 submetallicus White, Colpodes 11 suborbithorax Broun, Anchomenus 32 sulcitarsis Broun, Anchomenus 23 sulcitarsis (Broun), Kupeplatynus 23 takahe new species, Ctenognathus 44 taranaki new species, Kiwiplatynus 21 tawanui new species, Ctenognathus 42 tepaki new species, Ctenognathus 36 Tuiplatynus new genus 26 urewera new species, Ctenognathus 37 waimana new species, Prosphodrus 16 waltoni Britton, Prosphodrus 14 xanthomelus Broun, Anchomenus 51 xanthomelus (Broun), Ctenognathus 51 Zabronothus Broun 2