Taxonomic and nomenclatural history of Neckera (Bryophyta, Neckeraceae), including reinstatement of Rhystophyllum, the correct name for a segregate of this genus

59 Taxonomic and nomenclatural history of Neckera (Bryophyta, Neckeraceae), including reinstatement of Rhystophyllum, the correct name for a segregate of this genus Ryszard Ochyra1, Vítězslav Plášek2,3 , John C. Brinda4 1 National Biodiversity Collection, W. Szafer Institute of Botany, Polish Academy of Sciences, Lubicz 46, 31-512 Kraków, Poland 2 Department of Biology and Ecology, University of Ostrava, Chittussiho 10, 710 00 Ostrava, Czech Republic 3 InstituteofBiology,UniversityofOpole,Oleska48,PL‐45‐052Opole,Poland 4 Missouri Botanical Garden, 4344 Shaw Boulevard, Saint Louis, MO 63110, USA Correspondingauthor:VítězslavPlášek(Vitezslav[email protected]) Copyright: © Ryszard Ochyra et al. This is an open access article distributed under terms of the Creative Commons Attribution License (Attribution 4.0 International – CC BY 4.0). Research Article Abstract The complex taxonomic history of the moss genus Neckera, whose name was conserved at the Paris Congress in 1954, is reviewed. The issue of typifications of moss generic names by W. Ph. Schimper in 1860 is examined in detail and it is concluded that these typifications comply with the provisions of the "International Code of Nomenclature for algae, fungi and plants". Schimper was the first to typify Neckera with N. pennata, so it is unnecessary to treat this as a conserved type. The unitary designations of moss species proposed by Jakob Friedrich Ehrhart in his exsiccata "Phytophylacium Ehrhartianum", published between 1780 and 1785, are reviewed. It is concluded that two of these designations, Diphyscium and Paludella, are currently in use, having been validly published in the early nineteenth century. The same applies to Rhystophyllum, another of Ehrhart’s unitary designations, which was validated as a subsection of Neckera by C. Müller in 1850 and, subsequently, first elevated to the rank of section by Mitten in 1869, rather than Braithwaite in 1905 as listed in "Index muscorum". Finally, E. Britton raised this infrageneric taxon to the rank of genus in 1904. Rhystophyllum is here reinstated from obscurity, since it is homotypic with the generic name Exsertotheca that was recently introduced for a segregate of Neckera. Accordingly, three new combinations are proposed: Rhystophyllum crispum, Rh. intermedium and Rh. baeticum. Neckera subsect. Leiophyllum is regarded as a heterotypic synonym of Alleniella, another recent segregate of the formerly broadly circumscribed genus Neckera. Key words: Alleniella, Exsertotheca, generic names, infrageneric taxa, Musci, nomenclature, nomina generica conservanda, pleurocarpous mosses, Madrid Code, taxonomy, unitary designations of species Introduction The rapid advancement of phylogenetic taxonomy in recent decades has led to the increasing subdivision of large, traditionally recognised genera. These genera were originally circumscribed using methods of classical descriptive taxonomy, based primarily on morphological and anatomical features. As a result, many have since been split into multiple, smaller genera. The newly-delineated Academic editor: Matt von Konrat Received: 10 September 2025 Accepted: 10 November 2025 Published: 2 December 2025 Citation: Ochyra R, Plášek V, Brinda JC (2025) Taxonomic and nomenclatural history of Neckera (Bryophyta, Neckeraceae), including reinstatement of Rhystophyllum, the correct name for a segregate of this genus. PhytoKeys 267: 59–80. https://doi.org/10.3897/ phytokeys.267.171699 PhytoKeys 267: 59–80 (2025) DOI: 10.3897/phytokeys.267.171699 60 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera genera are typically assigned new names, a practice permitted under the "International Code of Nomenclature" (ICN; Turland et al. (2025)). However, it occasionally emerges that some generic names, validly published in the distant past, but subsequently forgotten, still meet all the necessary criteria for use and may be appropriately applied to these newly-recognised segregates. This article examines one such overlooked genus which, despite having fallen into obscurity, warrants reinstatement and application to one of the segregates of Neckera Hedw., a large and well-known genus of pleurocarpous mosses in the family Neckeraceae Schimp. Neckera has recently been subdivided into several smaller genera (Enroth 1991; Olsson et al. 2009a, 2009b, 2011, 2012, 2016; Enroth et al. 2022), for which new names have been proposed. One of these segregates, Exsertotheca S.Olsson, Enroth & D.Quandt, includes a common European calcicolous species previously known as N. crispa Hedw. This article reviews the taxonomic and nomenclatural history of the genus Neckera, with particular emphasis on its infrageneric classification. In many cases, the names of infrageneric taxa can be adopted for segregate genera through a simple change in status, thereby eliminating the need to coin entirely new names and to provide separate diagnoses and descriptions. Brief taxonomic history of Neckera From Hedwig to Müller Hallensis Neckera Hedw. is amongst the earliest generic names in muscology, first introduced by Hedwig (1782) under the orthographic variant Neckeria Hedw. (Fig. 1). The name commemorates Noël Martin Joseph de Necker (1730– 1793), a French-born botanist and bryologist who served as the personal physician to the court of the Electoral Palatinate in Mannheim, Germany (Frahm and Eggers 2001). Although Necker strongly opposed Hedwig’s view that mosses reproduce sexually, asserting instead that they do not (Schofield 1985), Hedwig nonetheless recognised Necker’s substantial contributions to botany, a fact he noted in justifying his choice of this eponym. Originally, four species were placed in Neckeria, including Hypnum crispum L. A decade later, Hedwig (1792) corrected the orthography of this name to Neckera and expanded the genus to include nine additional species, such as N. pennata Hedw. and N. pumila Hedw. However, neither Neckeria Hedw. (1782) nor Neckera Hedw. (1792) constitute validly published names under modern nomenclatural rules, as both appeared prior to 1801, the year Hedwig (1801) published "Species muscorum frondosorum". This work is the designated starting-point for moss nomenclature (excluding Sphagnaceae) under the ICN (Turland 2013; Turland et al. 2025), in contrast to most other groups whose nomenclature begins with Linnaeus (1753). Hedwig (1801) validated the generic name Neckera by giving a short diagnosis which included very general features of the peristome, namely that it is double and consists of 16 teeth of the outer peristome alternating with 16 distinct ciliate processes of the inner peristome (Fig. 2). The genus was quickly and widely accepted; however, under this broad circumscription, Neckera could encompass most pleurocarpous mosses that were primarily classified in the former order 61 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Isobryales (= Leucodontales). Indeed, this was the case, as throughout the past two centuries, over 600 specific and about 100 varietal epithets, as well as over 100 binary designations, were combined with Neckera (Wijk et al. 1964). One of the most striking examples of Neckera serving as a catch-all genus is found in the broad concept proposed by Müller (1850, 1851), published in September 1850 and supplemented in October 1851 (Stafleu and Cowan 1981). Müller assigned 199 species, known in the first half of the nineteenth century, to this single genus, subdividing them into nine sections and thirteen subsections, which highlighted its remarkable morphological heterogeneity. In modern classification systems, these species are now distributed across dozens of genera and multiple families of pleurocarpous mosses. In one case, the group even includes an acrocarpous species: Dicnemoloma pallidum (Hook.) Wijk & Margad. Of the 199 species included in Müller's (1850, 1851) synopsis of the genus, only 20 correspond to Neckera as currently circumscribed. Figure 1. The original publication of the generic name Neckeria Hedw. (from Hedwig 1782). Figure 2. The original publication of the generic name Neckera Hedw. (from Hedwig 1801). 62 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Schimper’s concept of Neckera and its development By a remarkable coincidence, just three months after publication of Müller’s (1850) concept of Neckera − in December 1850 − Schimper (1850a), in another monograph forming part of the monumental opus "Bryologia europaea" (Stafleu and Cowan 1976), presented a concept of this genus that was diametrically opposed to Müller’s. He assigned seven European species to the genus, including N. pennata, N. pumila, N. crispa, N. oligocarpa Bruch, N. complanata (Hedw.) Huebener, N. sendtneriana Schimp. and N. philippeana Schimp., all of which were described and illustrated in detail. In addition, he referenced five extra-European species: N. menziesii Hook. then known only from North America; N. hornschuchiana Müll. Hal. and N. angustifolia Müll.Hal. from Mexico; N. chilensis Schimp. from Chile; and N. jamesonii Taylor from Ecuador, but without comment. He also listed nine further designations in the protologue, including four from Africa, two each from Mexico and South America and one from Tasmania, all ascribed to W. Ph. Schimper. The species cited in the protologue roughly correspond to the two subsections distinguished by Müller (1850) within section ‘Euneckera’ Müll.Hal., i.e. the type section of Neckera, namely subsect. Leiophyllum Müll.Hal. and subsect. Rhystophyllum Müll.Hal. This section encompassed a number of species that are now classified under other genera, mostly related to Neckera or the Neckeraceae, such as Homaliodendron M.Fleisch., Porothamnium M.Fleisch., Neckeropsis Reichardt, Himantocladium (Mitt.) M.Fleisch., Calyptothecium Mitt. and Trachyloma Brid. Schimper’s (1850a) newly-proposed taxonomic concept of Neckera was quickly accepted in both Europe and North America, although the genus is not particularly species-rich in these regions. As exploration of tropical and subtropical areas advanced, new species of neckeroid mosses were discovered. These species often bore only superficial morphological similarities to Neckera, but due to the underdeveloped generic classification within the Neckeraceae, they were provisionally assigned to Neckera anyway (e.g. Mitten (1859, 1869); Dozy and Molkenboer (1863); Bescherelle (1872)). Consequently, many of these species were later transferred to other genera. In "Adumbratio florae muscorum totius orbis terrarum", published in the 1870s, Jaeger and Sauerbeck (1877, 1879) included 92 species and nine designations within Neckera. However, only 50 of those species could be retained in Neckera under the current taxonomic circumscription of this genus. Over the next quarter-century, the number of species assigned to Neckera grew to approximately 150. However, the number of species truly belonging to the genus in the strict sense increased to little more than a dozen (Paris 1896, 1900, 1905). Finally, Brotherus (1906) published the first global taxonomic overview of the genus Neckera in Engler’s "Die natürlichen Pflanzenfamilien" series, providing general keys to the species. He recognised 127 species within the genus, which were organised into six sections. This concept differed significantly from Schimper’s (1850a) circumscription of Neckera, as nearly half of the species were later found to belong to what are now widely accepted as separate genera − Neckeropsis and Himantocladium. These species were formally transferred to the respective genera in the second edition of the moss treatment in "Die natürlichen Pflanzenfamilien" (Brotherus 1925), reducing the number of Neckera species to 89. This taxonomic framework remained influential throughout the twentieth century, although the number of recognised species continued 63 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera to fluctuate. Names of many previously described species were synonymised with earlier ones, while others continued to be newly described. According to Crosby et al. (2000), Neckera comprised 71 species, with 27 of these remaining uninvestigated since their original description. Genus Neckera in the molecular era A new chapter in the taxonomic study of Neckera began over the past two decades, marked by the integration of phylogenetic taxonomic methods, based on molecular analyses (Olsson et al. 2009a, 2009b, 2010, 2011, 2012, 2016; Enroth et al. 2022). This advancement led to a re-definition of the genus, which was subsequently divided into three segregates. Amongst them are Alleniella S.Olsson, Enroth & D.Quandt, comprising 15 species and Exsertotheca S.Olsson, Enroth & D.Quandt, containing three species (Olsson et al. 2011; Enroth et al. 2022). In addition, six monotypic genera have been newly established or reinstated to accommodate certain Neckera species. These include Caduciella Enroth (Enroth 1991), Pengchengwua S.Olsson, Enroth, Huttunen & D.Quandt (Olsson et al. 2016), Enrothia Ignatov & Fedosov (Fedosov and Ignatov 2019), as well as Indoneckera Enroth, Taiwanobryopsis Enroth and Metaneckera Steere (Enroth et al. 2022). Furthermore, eight species have been transferred to the genus Taiwanobryum Nog. (Olsson et al. 2010; Ma et al. 2018; Enroth and Shevock 2021) and ten species have been re-assigned to Forsstroemia Lindb. (Olsson et al. 2011; Enroth et al. 2019, 2022). As a result of these taxonomic revisions, the genus Neckera currently comprises 39 extant and two fossil species (Brinda and Atwood 2025). Of the extant species, only 23 are widely recognised and accepted in taxonomic monographs, revisions and descriptive floras. The remaining 16 species are known solely from their original descriptions and have yet to undergo comprehensive taxonomic evaluation. Nomenclatural history of the generic name Neckera Conservation and former typifications The generic name Neckera has a rather complex nomenclatural history. When Hedwig (1782) first published the name Neckeria for a newly-established genus of mosses, it was predated by an identical name used by Scopoli (1777) for a flowering plant in the family Papaveraceae. Even when later the spelling of this name was corrected to Neckera (Hedwig 1792, 1801), it was, from a nomenclatural perspective, merely an orthographic variant and, thus, considered a later homonym of Neckeria Scop. As such, it could not be validly used as a name for an accepted moss genus. To resolve this issue, Neckera Hedw. was formally proposed for conservation prior to the Sixth International Botanical Congress held in Amsterdam in 1935 (Dixon 1934). While this proposal was recommended by the nomenclature committee at that Congress (Dixon 1935; Magill 1993) and Neckera was placed in the list of "Nomina generica conservanda" in Appendix V of the "Stockholm Code" (Lanjouw et al. 1952), it was not formally acted upon until much later because no bryophyte names were formally conserved prior to ratification at the Paris Congress in 1954 (see below). 64 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera The typification of a generic name is fundamental to the taxonomic interpretation of a genus. This is particularly critical for older names, which often encompassed a wide array of diverse and, sometimes, unrelated species. One such name is Neckera, whose typification also has had a complex and convoluted history. According to Crosby (1968), Grout (1934) was the first to designate N. crispa as the type. However, Grout (1939) later “amended” this designation in favour of N. pennata. As noted by Brinda and Davidse (2020), this change appears to have been influenced by discussions amongst members of the nomenclature committee, who selected N. pennata as the type instead (Dixon 1939). However, this decision conflicts with Art. 10.5 of the "Code" (Turland et al. 2025), which would give precedence to the earlier designation of N. crispa. Nevertheless, the revised typification was accepted by the General Committee and Advisory Board at the Eighth International Botanical Congress in Paris in 1954. Since the publication of the "Paris Code" (Anonymous 1954; Lanjouw et al. 1956), N. pennata has been listed as the type species of Neckera. Considering Grout’s earlier designation of N. crispa in 1934, N. pennata has been included in Appendix III of the "Code" as a conserved type since the "Montreal Code" (Lanjouw et al. 1961). Crosby (1968) opposed this outcome and proposed reinstating N. crispa as the generitype of Neckera, but his proposal was ultimately rejected by the Committee for Bryophytes (Florschütz 1973). Consequently, the recognition of N. pennata as a conserved type has achieved broad consensus (Wijk et al. 1964; Farr et al. 1979a; Greuter et al. 1993). Schimper’s typifications of moss generic names In accordance with the delimitation of Neckera in "Bryologia europaea" (Schimper 1850a) and its subsequent usage, N. pennata Hedw. was formally designated as the type of this generic name by Schimper (1860) in the first edition of his "Synopsis muscorum europaeorum". This typification has largely been overlooked by bryologists, despite the relevant information appearing verbatim on the title page of the work: “Accedunt tabulae VIII typos genericos exhibentes, ...” [= “There are also eight plates showing generitypes” (emphasis ours)]. This critical phrase was first identified and interpreted in the mid-1990s by W. D. Margadant, who prepared a manuscript on Schimper’s typifications, which was submitted to Lindbergia. Pekka Isoviita of Helsinki was invited to serve as a referee for the manuscript and undertook a meticulous review, introducing numerous corrections and amendments. As a result, Margadant proposed that Isoviita be included as a co-author of the paper (Isoviita, personal communication to R. Ochyra in litteris, 23 February 1995; Margadant and Geissler (1995)). Unfortunately, the planned publication was never completed and published. However, typifications for four generic names, Dicranella (Müll.Hal.) Schimp., Hylocomium Schimp., Pylaisia Schimp. and Homalia Brid., were later published by Margadant and Geissler (1995) as part of formal proposals to conserve these names. For three of these, conservation of the types was necessary to maintain current usage, which had been threatened by Schimper’s (1860) overlooked type designations involving different species. In the case of Homalia, however, the typification of the name was merely confirmed by Schimper (1860), since in "Bryologia europaea", Schimper (1850b) had used the term Grundform, in the German text only, as an equivalent of “type”. All these proposals were accepted 65 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera by the Committee for Bryophyta (Zijlstra 1998, 1999) and the conserved generic names were subsequently included in Appendix III of the "Saint Louis Code" (Greuter et al. 2000), thereby formally endorsing Schimper’s (1860) typifications. In addition to the article by Margadant and Geissler (1995), published in the November issue of Taxon, Volume 44, Margadant’s discovery was also acknowledged in the monograph on the genus Racomitrium Brid. in Poland (Bednarek-Ochyra 1995). Notably, this monograph was published even earlier, on 1 October 1995, than the aforementioned article. As mentioned above, the information regarding the typification of generic names in Schimper’s (1860) work had been communicated to R. Ochyra by P. Isoviita, who also granted permission for its inclusion in the monograph of Racomitrium. This case is of particular interest because Schimper (1860) designated R. lanuginosum (Hedw.) Brid. as the type, thereby rendering superfluous a later typification by Pfeiffer (1874), who had proposed R. canescens (Hedw.) Brid. instead. This change in type somewhat affected the nomenclature of infrageneric taxa within Racomitrium and subsequently influenced the naming of segregate genera derived from this heterogeneous and broadly circumscribed genus (Bednarek-Ochyra et al. 2014; Sawicki et al. 2015). For the next quarter-century, bryologists largely overlooked the typification of generic names in Schimper’s (1860) opus. Zijlstra (2015) reignited the debate by claiming that the phrase on the book’s title page implies the figures represent typical species. This, however, is a clear overstatement: Schimper (1860) explicitly used the term type (typus), not typical (typicus). While it is possible that Schimper meant “exemplar”, not our modern nomenclatural type, that is irrelevant under the current wording of the "Code" (Turland et al. 2025). Notably, the same terminology appears in "Nomenclator botanicus" by Pfeiffer (1873a), who included a general statement on the unnumbered second page of the Prefatio: “Species plantarum in libro meo omnino negliguntur, excepta indicatione illarum, quae typum generis novi aut novo modo circumscripti vel sectionis offerunt” [= “Plant species are entirely neglected in my book, except for indication of those which are presented as the type of a new or re-circumscribed genus or of a section” (emphasis ours)]. Like Schimper (1860), Pfeiffer (1873a) clearly indicated type-species for generic or sectional names and these typifications are acceptable under the "Code" (Art. 7.11, Ex. 18; Turland et al. (2025)). Of course, it occasionally emerges that Schimper’s or Pfeiffer’s choices were erroneous or based on a misinterpretation of the protologue. However, such cases do not undermine the fundamental validity of their typifications. Similar issues occur even in contemporary practice and erroneous choices can be corrected in accordance with Art. 10 of the "Code" (Turland et al. 2025). Typification of moss generic names by Schimper (1860) once again drew the attention of the Committee for Bryophytes in the early 2020s, during discussions surrounding the proposal to conserve the name Oreas Brid. (Brinda and Fedosov 2020). Although the name was formally proposed for conservation with the conserved type Oreas martiana (Hoppe & Hornsch.) Brid., based on the then-prevailing assumption that Oreas had first been typified by Pfeiffer (1873b) with “Weissia Mielichhoferi Schwägr.” (≡ Weissia mielichhoferiana Funck), the Committee ultimately accepted Schimper’s (1860) earlier typification of this generic name with O. martiana as effectively published (Klazenga 2023). This decision was subsequently accepted by the General Committee with the suggestion that “bryologists should check whether this acceptance of 66 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Schimper’s typification in his 1860 paper has consequences for typifications of other names in that paper” (Wilson 2023). However, most of the issues arising from Schimper’s (1860) typifications, which had significantly affected the status of several of the aforementioned generic names, have in fact already been resolved via conservation proposals (Margadant and Geissler 1995). Schimper’s (1860) typifications have recently gained broader recognition, as reflected in the TROPICOS database (https://www.tropicos.org/), where an attempt has been made to correctly list the types of all bryophyte genera. This includes the generic name Neckera, which was first typified with N. pennata (https://www.tropicos.org/name/35000848). In Table VI, Schimper (1860) presented an illustration labelled only with the name Neckera. However, the accompanying legend specifies N. pennata and includes captions identifying the illustrated details. Consequently, Dixon’s (1939) seemingly arbitrary selection of N. pennata aligns with Schimper’s earlier, but long-overlooked typification. This finding clarifies that Dixon’s (1939) choice was indeed correct and that Grout’s (1934) typification was not the earliest. Therefore, N. pennata should not be listed as a conserved type in Appendix III of the "Code", as suggested by Brinda and Davidse (2020), but rather its current status should be maintained. As noted above, N. pennata was treated as the conserved type of Neckera from the "Montreal Code" (Lanjouw et al. 1961) through to the "Tokyo Code" (Greuter et al. 1994). However, in the mid-1990s, W. D. Margadant re-discovered Schimper’s (1860) earlier typifications of many generic names, including Neckera. As a result, treating N. pennata as a conserved type became unnecessary. Accordingly, beginning with the "Saint Louis Code" (Greuter et al. 2000), the phrase "typus conservandus" associated with N. pennata was editorially removed. Synonyms of the generic name Neckera In summary, the generic name Neckera currently has two heterotypic synonyms and ten homotypic synonyms (Brinda and Atwood 2025). The first heterotypic synonym is Alsia Sull., formerly treated as a monotypic genus endemic to the Pacific coast of North America (Schofield 1980). It included only A. californica (Hook. & Arn.) Sull., a species originally described as Neckera californica Hook. & Arn., which has recently been transferred back to Neckera (Olsson et al. 2011). The second is Neckera sect. Douglasiella (Kindb.) Paris, typified by the North American species N. douglasii Hook. Three of the homotypic synonyms are homonyms and seven represent names of homotypic infrageneric taxa. Of these, Distichia (Brid.) Brid. and Braunia Hornsch. are considered nomina rejicienda, while Cryptopodium Fürnr. is both a homonym and a superfluous name by typification. Additionally, Cryptopodia (Röhl.) Fürnr. is a superfluous name by typification and Eleutera P.Beauv. ex Stuntz is illegitimate, as it included the previously designated type of Neckera. Taxonomic and nomenclatural history of Rhystophyllum Earliest plant exsiccatae The name Rhystophyllum first appeared in bryological literature through the work of Jakob Friedrich Ehrhart (1780–1785, 1789), who used it on the label of a specimen distributed in 1780 as No. 79 in the eighth decade (a set of ten 67 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera numbered specimens) of his exsiccata "Phytophylacium Ehrhartianum" (Fig. 3). Ehrhart (1742–1795), a German botanist of Swiss origin and a gifted student of Linnaeus, held the position of Royal Botanist in the Electorate of Hanover (Braunschweig-Lüneburg) during the final decade of his life. Ehrhart is best known in the history of botany as one of the pioneers in publishing exsiccatae, numbered collections of dried and pressed plant specimens, distributed under a common title, precisely identified and accompanied by printed labels, intended for circulation amongst botanists and institutions. Although Johann Balthasar Ehrhart (1700–1756) is regarded as the first publisher of an exsiccata, his "Herbarium vivum recens collectum", issued in 1732, contained only medicinal plants and no cryptogams. Today, only three copies of this earliest exsiccata are known to exist (Stafleu 1972). On the other hand, Jakob Friedrich Ehrhart – the similarity of names is coincidental – published for commercial purposes seven series of exsiccatae in the years 1780–1793, covering about 1620 species of flowering and cryptogamic plants (Britten 1922, 1923; Stafleu and Cowan 1976; Sokoloff et al. 2002). For bryologists, the first two series are particularly important, the above-mentioned "Phytophylacium Ehrhartianum" and "Plantae cryptogamae Linneae" (Sayre 1969), comprising 320 specimens of mosses, liverworts, lichens, algae, club-mosses, horsetails and ferns, distributed in decades (Ehrhart 1785–1793, 1792). In the first of these series, only six species of mosses and ten lichens were distributed from the cryptogams. However, they are particularly interesting from a nomenclature Figure 3. The specimen No. 79 from "Phytophylacium Ehrhartianum" from UPS with the unitary designation Rhystophyllum of Neckera crispa Hedw. 74 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Brotherus VF (1906) Neckeraceae. In: Engler A, Prantl K (Eds) Die Naturlichen Pflanzenfamilien nebst ihren Gattungen und wichtigeren Arten inbesondere den Nutzpflanzen. I. Teil. 3. Abteilung. II. Hälfte: Musci (Laubmoose) III. Unterklasse Bryales: II. Spezieller Teil. II. Gruppe: Pleurocarpi. Verlag von Wilhelm Engelmann, Leipzig, 775−863. Brotherus VF (1925) Neckeraceae. In: Engler A (Ed.) Die Naturlichen Pflanzenfamilien nebst ihren Gattungen und wichtigeren Arten inbesondere den Nutzpflanzen. 11. Band. Musci (Laubmoose) 2. Hälfte. Verlag von Wilhelm Engelmann, Leipzig, 178−202. Crosby MR (1968) (241) Proposal to change the listed type of the genus Neckera Hedw. (nom. cons.), Musci. Taxon 17(5): 594–595. https://doi.org/10.2307/1216079 Crosby MR, Magill RE, Allen BH, He S (2000) A checklist of the mosses. Missouri Botanical Garden, St. Louis, vi + 320 pp. Dixon HN (1934) [1935] Bryological nomenclature. Revue Bryologique et Lichénologique. Nouvelle Série 7 (3−4): 137–141. Dixon HN (1935) Bryology at the sixth international Congress. Revue Bryologique et Lichénologique, Nouvelle Série 8(3−4): 226. Dixon HN (1939) International bryological nomenclature. The Journal of Botany. British and Foreign 77: 176–178. Dozy F, Molkenboer JH (1863) Bryologia javanica seu descriptio muscorum frondosorum Archipelagi indici iconibus illustrata. Vol. 2, Fasc. 37−40. E. J. Brill, Lugduni-Batavorum, 57−72. [+ Tab. clxxxi−cc] Ehrhart F (1779) Webera, eine Pflanzengattung. Hannoverisches Magazin 17: columns 257−258. Ehrhart F (1789) Index Phytophylacii Ehrhartiani. In: Beiträge zur Naturkunde und den damit verwandten Wissenschaften, besonders der Botanik, Chemie, Hausund Landwirtschaft, Arzneigelahrtheit und Apothekerkunst. Vierter Band. Im Verlage der Schmidtischen Buchhandlung, Hannover und Osnabrück, 145−150. Ehrhart F (1792) Index plantarum cryptogamarum Linn., quas in locis earum natalibus collegit et exsicavit Fridericus Ehrhart, Helveto-Bernas. In: Beiträge zur Naturkunde und den damit verwandten Wissenschaften, besonders der Botanik, Chemie, Hausund Landwirtschaft, Arzneigelahrtheit und Apothekerkunst. Siebenter Band. Im Verlage bei Christian Ritscher, Hannover und Osnabrück, 94−102. Ehrhart F (1780–1785) Phytophylacium Ehrhartianum, continens plantas quas in locis earum natalibus collegit et exsiccavit Fridericus Ehrhart, Helveto-Bernas. Decades I–X, Numeri 1−100. Hannoverae. Ehrhart F (1785–1793) Plantae cryptogamae Linneae, quas in locis earum natalibus collegit et exsicavit Fridericus Ehrhart, Helveto-Bernas. Decades I–XXXII, Numeri 1−320. Sacra, Hannoverae. Enroth J (1991) Notes on the Neckeraceae (Musci). 10. The taxonomic relationships of Pinnatella mairei, with the description of Caduciella (Leptodontaceae). Journal of Bryology 16(4): 611–618. https://doi.org/10.1179/jbr.1991.16.4.611 Enroth J, Shevock JR (2021) Contributions to the moss families Neckeraceae and Pterobryaceae of Bhutan. Arctoa 30(1): 43–46. https://doi.org/10.15298/arctoa.30.04 Enroth J, Fedosov VE, Fedorova AV, Ignatova EA, Ignatov MS (2019) Miscellaneous notes on the genus Forsstroemia (Neckeraceae, Bryophyta) in Russia. Arctoa 28(1): 18–23. https://doi.org/10.15298/arctoa.28.03 Enroth J, Olsson S, Huttunen S, Quandt D (2022) Neckera, Forsstroemia and Alleniella (Neckeraceae, Bryophyta) redefined based on phylogenetic analyses. The Bryologist 125(2): 311–327. https://doi.org/10.1639/0007-2745-125.2.311 75 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Fabricius PhC (1743) Primitiae florae butisbacensis, sive sex decades plantarum rariorum inter alias circa Butisbacum sponte nascentium, cum observationibus methodos plantarum Turnefortianam, Rivinianam, Raianam, Knautianam, et Linnaeanam potissimum concernentibus, recensitae, et celeberrimo nomini viri excellentissimi et experientissimi domini D. Iohan. Casimiri Hertii, consiliarii et archiatri Hasso-Darmstadini medicinaeque in Alma Giessena Professoris Primarii, inscriptae. Literis Wincklerianis, Wetzlariae, 64 pp. Farr ER, Leussink JA, Stafleu FA [Eds] (1979a) Index nominum genericorum (plantarum). Volume 2. Eprolithus – Peersia. Regnum Vegetabile 101. Bohn, Scheltema & Holkema. Utrecht and dr. W. Junk b.v., Publishers, The Hague, 631–1276. Farr ER, Leussink JA, Stafleu FA [Eds] (1979b) Index nominum genericorum (plantarum). Volume 3. Pegaeophyton – Zyzygium. Regnum Vegetabile 102. Bohn, Scheltema & Holkema. Utrecht and dr. W. Junk b.v., Publishers, The Hague, 1277–1896. Fedosov VE, Ignatov MS (2019) Enrothia, a new genus of Neckeraceae (Bryophyta) from East Asia. Arctoa 28(1): 12–17. https://doi.org/10.15298/arctoa.28.02 Florschütz PA (1973) [Report of] the Committee for Bryophytes. Taxon 22(1): 157–158. https://doi.org/10.2307/1218068 Frahm J-P, Eggers J (2001) Lexikon deutschsprachiger Bryologen. Books on Demand GmbH (BoD), Norderstedt, 672 pp. Greuter W, Brummitt RK, Farr E, Kilian N, Kirk PM, Silva PC (1993) NCU–3. Names in current use for extant plant genera. Regnum Vegetabile 129. Koeltz Scientific Books, Königstein, xxvii + 1464 pp. Greuter W, Barrie FR, Burdet HM, Chaloner WG, Demoulin V, Hawksworth DL, Jørgensen PM, Nicolson DH, Silva PC, Trehane P, McNeill J [Eds] (1994) International Code of Botanical Nomenclature (Tokyo Code) adopted by the Fifteenth International Botanical Congress, Yokohama, August–September, 1993. Regnum Vegetabile 131. Koeltz Scientific Books, Königstein, xviii + 389 pp. Greuter W, McNeill J, Barrie FR, Burdet HM, Demoulin V, Filgueiras TS, Nicolson DH, Silva PC, Skog JE, Trehane P, Turland NJ [Eds] (2000) International Code of Botanical Nomenclature (Saint Louis Code) adopted by the Sixteenth International Botanical Congress, Saint Louis, Missouri, July–August, 1999. Regnum Vegetabile 138. Koeltz Scientific Books, Königstein, xviii + 474 pp. Grout AJ (1934) Family Neckeraceae. In: Moss flora of North America north of Mexico, Vol. 1, Part 4. Published by the author, Newfane, Vermont, 248−253. Grout AJ (1939) Additions and corrections. In: Moss flora of North America north of Mexico, Vol. 3, Part 4. Published by the author, Newfane, Vermont, 208−213. [+ pl. 65, figs 1−10 (left)] Haller A (1742) Enumeratio methodica stirpium Helvetiae indigenarum. Qua omnium brevis descriptio et synonymia compendium virium medicarum dubiarum declaratio novarum et rariorum uberior historia et icones continentur. Tomus 1. Ex Officina Academica Abrami Vandenhoek, Gottingae, i−iv + 1−36 (Prefatio) + 1−424. Hartman C (1871) Handbok i Skandinaviens flora, innefattande Sveriges och Norges växter, till och med mossorna. Tionde upplagan, utoifven med rättesler och tillägg. Sednare delen: mossor. Zacharias Haeggströms förlag, Stockholm, xxviii + 180 pp. Hedwig J (1782) Fundamentum historiae naturalis muscorum frondosorum concernens eorum flores, fructus, seminalem propagationem adiecta generum dispositione methodica, iconibus illustratis. Pars II. Apud Siegfried Lebrecht Crusium, Lipsiae, xii + 108 pp. [+ 10 tab.] https://doi.org/10.5962/bhl.title.82187 76 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Hedwig J (1792) Descriptio et adumbratio microscopico-analytica muscorum frondosorum novorum dubiisque vexatorum. Volumen tertium. In Bibliopolio Gleditschiano, Lipsiae, 100 pp. [+ 40 tab.] Hedwig J (1801) Species muscorum frondosorum descriptae et tabulis aeneis LXXVII coloratis illustratae. Sumtu Joannis Ambrosii Barthii & apud Amand Koenig, Parisiis, vi + 353 pp. [+ lxxvii tab.] https://doi.org/10.5962/bhl.title.26 Hedwig J (1785−1787) Descriptio et adumbratio microscopico-analytica muscorum frondosorum novorum dubiisque vexatorum. Volumen primum. In Bibliopolio Gleditschiano, Lipsiae, xviii + 110 pp. [+ 40 tab.] https://doi.org/10.5962/bhl.title.705 Jaeger A, Sauerbeck Fr (1877) Genera et species muscorum systematyce disposita seu adumbratio florae muscorum totius orbis terrarum. Pars 7. Bericht über die Thätigkeit der St. Gallischen Naturwissenschaftlichen Gesellschaft während des Vereinjahres 1875−76 [Band 17]: 201−371. Jaeger A, Sauerbeck Fr (1879) Genera et species muscorum systematyce disposita seu adumbratio florae muscorum totius orbis terrarum. Pars 9. Bericht über die Thätigkeit der St. Gallischen Naturwissenschaftlichen Gesellschaft während des Vereinjahres 1877−78 [Band 19]: 257−514. Klazenga N (2023) Report of the Nomenclature Committee for Bryophytes: 14. Taxon 72(3): 654–655. https://doi.org/10.1002/tax.12940 Lanjouw J, Baehni Ch, Merrill ED, Rickett HW, Robyns W, Sprague TA, Stafleu FA [Eds] (1952) International Code of Botanical Nomenclature adopted by the Seventh International Botanical Congress, Stockholm, July, 1950. Regnum Vegetabile 3. International Bureau for Plant Taxonomy and Nomenclature of the International Association for Plant Taxonomy, Utrecht – Netherlands, 228 pp. Lanjouw J, Baehni Ch, Robyns W, Rollins RC, Ross R, Rousseau J, Schulze GM, Smith AC, de Vilmorin R, Stafleu FA [Eds] (1956) International Code of Botanical Nomenclature adopted by the Eighth International Botanical Congress, Paris, July, 1954. Regnum Vegetabile 8. International Bureau for Plant Taxonomy and Nomenclature of the International Association for Plant Taxonomy, Utrecht – Netherlands, 338 pp. Lanjouw J, Baehni Ch, Robyns W, Ross R, Rousseau J, Schopf JM, Schulze GM, Smith AC, de Vilmorin R, Stafleu FA [Eds] (1961) International Code of Botanical Nomenclature adopted by the Ninth International Botanical Congress, Montreal, August, 1959. Regnum Vegetabile 23. International Bureau for Plant Taxonomy and Nomenclature of the International Association for Plant Taxonomy, Utrecht – Netherlands, 372 pp. Limpricht KG (1894) Die Laubmoose Deutschlands, Oesterreichs und der Schweiz. I. Abtheilung: Sphagnaceae, Andreaeaceae, Archidiaceae, Bryineaqe (Cleistocarpae, Stegocarpae [Acrocarpae]. Lieferung 24. In: Dr. L. Rabenhorst’s Kryptogamen-Flora von Deutschlands, Oesterreichs und der Schweiz. Zweite Auflage. Band 4: Die Laubmoose Deutschlands, Oesterreichs und der Schweiz. Verlag von Eduard Kummer, Leipzig, 641−704. Lindberg SO (1879) Musci scandinavici in systemate novo naturali dispositi. Ex Officina Iesaiae Edquist, Upsaliae, 50 pp. Linnaeus C (1753) Species plantarum, exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundam systema sexuale digestas. Tomus 1−2. Impensiis Laurentii Salvii, Holmiae, xii + 1200. [+ 1−30, index] https://doi.org/10.5962/bhl.title.669 Linnaeus C (1760) Amoenitates academicae; seu dissertationes variae physicae, medicae, antehac seorsim editae, nunc collectae et auctae, cum tabulis aeneis. Volumen quintum. Sumtu & Litteris Direct. Laurentii Salvii, Holmiae, iv + 483 + [484−486, index dissertationum, 1−5, catalogus] [+ Tab. i−iii.] 77 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Linnaeus C (1763) Species plantarum, exhibentes plantas rite cognitas, ad genera relatas, cum differentiis specificis, nominibus trivialibus, synonymis selectis, locis natalibus, secundam systema sexuale digestas. Tomus 2. Editio secunda, aucta. Impensiis Direct. Laurentii Salvii, Holmiae, 785−1684. [+ 1−64, indexes] https://doi. org/10.5962/bhl.title.65726 Ma W-Z, Enroth J, Shevock JR (2018) Taiwanobryum undulatifolium (Neckeraceae), a new combination based on sporophytic features. Journal of Bryology 40(1): 51–55. https://doi.org/10.1080/03736687.2017.1409326 Magill RE (1993) Conserved names for mosses: A brief history. Taxon 42(1): 5–15. https://doi.org/10.2307/1223298 Margadant WD, Geissler P (1995) (1175−1191) Seventeen proposals concerning nomina generica conservanda for genera of Musci. Taxon 44(4): 613–624. https://doi. org/10.2307/1223506 Mitten W (1859) Musci Indiae Orientalis; an enumeration of the mosses of the East Indies. Journal of the Proceedings of the Linnean Society. Supplement to Botany 1: 1−171. https://doi.org/10.5962/bhl.title.156377 Mitten G (1869) Musci austro-americani. Enumeratio muscorum omnium austro-americanorum auctori hucusque cognitorum. The Journal of the Linnean Society Botany 12: 1–659. https://doi.org/10.1111/j.1095-8339.1871.tb00633.x Mohr DMH (1803) Observationes botanicae quibus consentiente Ampliss. Philosophor. Ord. Kilon. pro gradu Doctoris Philosophiae nec non LL. AA. Magistri rite obtinendo plantarum cryptogamarum ordines, genera et species ilustrare conatus est auctor Dan. Matth. Henr. Mohr, Quickbornâ-Pinnebergensis, Soc. Phys. Goetting. adser. Typis C. F. Mohr, Kiliae, 45 + [2] pp. https://doi.org/10.5962/bhl.title.53607 Müller C (1848) Synopsis muscorum frondosoum omnium hucusque cognitorum. Pars secunda. Musci vegetationis pleurocarpicae. Fasc. 1. Sumptibus Alb. Foerstner, Berolini, 1−160. https://doi.org/10.5962/bhl.title.31 Müller C (1850) Synopsis muscorum frondosoum omnium hucusque cognitorum. Pars secunda. Musci vegetationis pleurocarpicae. Fasc. 6. Sumptibus Alb. Foerstner, Berolini, 1−160. https://doi.org/10.5962/bhl.title.31 Müller C (1851) Synopsis muscorum frondosoum omnium hucusque cognitorum. Pars secunda. Musci vegetationis pleurocarpicae. Fasc. 9−10. Supplementum ad Synopsin muscorum frondosorum una cum conspectu systematis. Sumptibus Alb. Foerstner, Berolini, 511−772. Ochyra R (2001) The first valid publication of the Linnaean Mnium triquetrum (Musci) and other new names for mosses proposed by N. Jolyclerc. Cryptogamie. Bryologie 22(1): 29–39. https://doi.org/10.1016/S1290-0796(00)01051-8 Ochyra R, Szmajda P, Bocheński W, Karczmarz K (1988) M. 451. Neckera crispa Hedw. In: Ochyra R, Szmajda P (Eds) Atlas of the geographical distribution of spore plants in Poland. Series 5. Mosses (Musci). Part 4. Państwowe Wydawnictwo Naukowe, Warszawa − Poznań, 27−31. [+ 1 map (folio)] [In Polish and English] Olsson S, Buchbender V, Enroth J, Hedenäs L, Huttunen S, Quandt D (2009a) Phylogenetic analyses reveal high levels of polyphyly among pleurocarpous lineages as well as novel clades. The Bryologist 112(3): 447–466. https://doi.org/10.1639/0007-2745-112.3.447 Olsson S, Buchbender V, Enroth J, Hedenäs L, Huttunen S, Quandt D (2009b) Evolution of the Neckeraceae (Bryophyta): Resolving the backbone phylogeny. Systematics and Biodiversity 7(4): 419–432. https://doi.org/10.1017/S1477200009990132 Olsson S, Buchbender V, Enroth J, Hedenäs L, Huttunen S, Quandt D (2010) Phylogenetic relationships in the “Pinnatella” clade of the moss family Neckeraceae (Bryophyta). 78 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Organisms, Diversity & Evolution 10(2): 107–122. https://doi.org/10.1007/s13127010-0017-z Olsson S, Enroth J, Buchbender V, Hedenäs L, Huttunen S, Quandt D (2011) Neckera and Thamnobryum (Neckeraceae, Bryopsida): Paraphyletic assemblages. Taxon 60(1): 36–50. https://doi.org/10.1002/tax.601005 Olsson S, Enroth J, Huttunen S, Quandt D (2012) Forsstroemia Lindb. (Neckeraceae) revisited. Journal of Bryology 34(2): 114–122. https://doi.org/10.1179/174328201 1Y.0000000047 Olsson S, Enroth J, Huttunen S, Quandt D (2016) Phylogeny of Neckeropsis and Himantocladium (Neckeraceae, Bryophytina). Bryophyte Diversity and Evolution 38(2): 53–70. https://doi.org/10.11646/bde.38.2.4 Paris (1896) Index bryologicus sive enumeratio muscorum hucusque cognitorum adjunctis synonymia distributioneque geographica locupletissimis. Pars 3. Apud Paul Klincksieck, Parisiis, 645−964. Paris (1900) Index bryologicus sive enumeratio muscorum hucusque cognitorum adjunctis synonymia distributioneque geographica locupletissimis. Supplementum primum. Georg et Cie, Libraires−Éditeurs, Genève et Bale; même maison, Lyon, 334 pp. Paris (1905) Index bryologicus sive enumeratio muscorum hucusque cognitorum adjunctis synonymia distributioneque geographica locupletissimis. Edition secunda. Vol. 3, Fasc. 17. Librairie scientifique A. Hermann, 265−328. Pfeiffer LKG (1873a) Nomenclator botanicus. Nominum ad finem anni 1858 publici juris factorum, classes, ordines, tribus, familias, divisiones, genera, subgenera vel sectiones designantium enumeratio alphabetica. Adjectis auctoribus, temporibus, locis systematicis apud varios, notis literariis atque etymologicis et synonymis. Vol. 1. Pars prior. Sumptibus Theodori Fischeri, Cassellis, i−vi + 1−808. https://doi.org/10.5962/ bhl.title.10853 Pfeiffer LKG (1873b) Nomenclator botanicus. Nominum ad finem anni 1858 publici juris factorum, classes, ordines, tribus, familias, divisiones, genera, subgenera vel sectiones designantium enumeratio alphabetica. Adjectis auctoribus, temporibus, locis systematicis apud varios, notis literariis atque etymologicis et synonymis. Vol. 1. Pars altera. Sumptibus Theodori Fischeri, Cassellis, 809−1876. https://doi.org/10.5962/bhl.title.10853 Pfeiffer LKG (1874) Nomenclator botanicus. Nominum ad finem anni 1858 publici juris factorum, classes, ordines, tribus, familias, divisiones, genera, subgenera vel sectiones designantium enumeratio alphabetica. Adjectis auctoribus, temporibus, locis systematicis apud varios, notis literariis atque etymologicis et synonymis. Vol. 2. Pars altera. Sumptibus Theodori Fischeri, Cassellis, 761−1698. https://doi.org/10.5962/ bhl.title.10853 Röhling JCh (1800) Moosgeschichte Deutschlands. Erster Theil. Die Beschreibung aller in Deutschland entdeckten Moosarten enthaltend. Bey Friedrich Wilmans, Bremen, xli + 436 pp. Röhling JCh (1808) Anhang. Systematisches Verzeichniss der auf dem Braubacher Gebiete gesammelten Laubmoose. In: Borkhausen’s Ringen nach dem schönsten Ziele des Mannes. Ein Denkmal der Freundschaft. Bey Friedrich Wilmans, Frankfurt am Mayn, 125−162. Röhling JCh (1813) Deutschlands Flora. Ein botanisches Taschenbuch. Dritter Theil. Kryptogamische Gewächse. Zweite, durchaus umgearbeitete Ausgabe, x + 210 pp. Sawicki J, Szczecińska M, Bednarek-Ochyra H, Ochyra R (2015) Mitochondrial phylogenomics supports splitting the traditionally conceived genus Racomitrium (Bryophyta: 79 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Grimmiaceae). Nova Hedwigia 100(3–4): 293–317. https://doi.org/10.1127/nova_ hedwigia/2015/0248 Sayre G (1969) Cryptogamae exsiccatae — An annotated bibliography of published exsiccatae of Algae, Lichenes, Hepaticae, and Musci. Memoirs of the New New York Botanical Garden 19(1): 1–174. Schimper WPh (1850a) Neckera. In: Schimper WPh (Ed.) Bryologia europaea seu genera muscorum europaeorum monographicae illustrata (Vol. 5). Sumptibus Librariae E. Schweizerbart, Stuttgartiae, 41−51. [+ Tab. 440−445] [Fasc. 44–45 Mon.: pp. 1−11 + Tab. 1−6]. Schimper WPh (1850b) Omalia. In: Schimper WPh (Ed.) Bryologia europaea seu genera muscorum europaeorum monographicae illustrata (Vol. 5). Sumptibus Librariae E. Schweizerbart, Stuttgartiae, 53−55. [+ Tab. 446] [Fasc. 44–45 Mon.: pp. 1−3 + Tab. 1] Schimper WPh (1860) Synopsis muscorum europaeorum praemissa introduction de elementis bryologicis tractante. Sumptibus Librariae E. Schweizerbart, Stuttgart, vi + lix + 733 pp. [+ 1 map + pls. 1−8] Schmidel CCh (1758) Dissertatio inauguralis botanica de Buxbaumia. Quam rectore magnificentissimo serenissimo ac principe domino domino Friderico [marggravio Brandenburgico Borussiae ac Silesiae caet. caet. Duce burggravio Norico reliqua rel. circuli Franconici duce militari supremo et trium legionem atque cohortium praefecto] ex decreto illustris et gratiosi ordinis medici praeside D. Casimiro Christophoro Schmidelio sereniss. marggr. Brand. Culmb. a consil. aul. med. anat. et botan. p. p. o. Acad. Imperial. Nat. Cur. et Societat. Botan. Florentinae sodali facult. Med. h. t. decano] in Alma Fridericiana Erlangensi pro doctoris gradu honoribus privilegiis ac immunitatibus rite obtinendis ad diem Jun. MDCCLVIII. Placidae eruditorum disquisition subiicit Iohannes Georgius Hoelzel wonsideliensis. Litteris Tetschnerianis, Erlangae, 40 pp. [+ Tab. i−ii (one plate)] Schofield WB (1980) Phytogeography of the mosses of North America (north of Mexico). In: Taylor RJ, Leviton AE (Eds) The mosses of North America. Pacific Division of the American Association for the Advancement of Science, San Francisco, 131−170. Schofield WB (1985) Introduction to bryology. Macmillan Publishing Company, New York & Collier Macmillan Publishers, London, xvi + 431 pp. Scopoli AJ (1777) Introductio ad historiam naturalem sistens genera lapidum, plantarum, et animalium hactenus detecte, caracteribus essentialibus donata, in tribus divisa, subinde ad leges naturae. Apud Wolfgangum Gerle, Bibliopolam, Pragae, x + 540 pp. https://doi.org/10.5962/bhl.title.10827 Sokoloff DD, Balandin SA, Gubanov IA, Jarvis CE, Majorov SR, Simonov SS (2002) The history of botany in Moscow and Russia in the 18th and early 19th centuries in the context of the Linnaean ollection at Moscow University (MW). Huntia 11(2): 129–191. Stafleu FA (1972) Cryptogamae exsiccatae. Taxon 21(2–3): 351–354. https://doi. org/10.2307/1218211 Stafleu FA, Cowan RS (1976) Taxonomic literature. A selective guide to botanical publications and collections with dates, commentaries and types. Volume 1: A−G. Ed. 2. Regnum Vegetabile 94. Bohn, Scheltema & Holkema, Utrecht, xl + 1136 pp. https:// doi.org/10.5962/t.206090 Stafleu FA, Cowan RS (1981) Taxonomic literature. A selective guide to botanical publications and collections with dates, commentaries and types. Volume 3: Lh−O. Ed. 2. Regnum Vegetabile 105. Bohn, Scheltema & Holkema. Utrecht and dr. W. Junk b.v., Publishers, The Hague, xii + 980 pp. https://doi.org/10.5962/t.207599 80 PhytoKeys 267: 59–80 (2025), DOI: 10.3897/phytokeys.267.171699 Ryszard Ochyra et al.: Rhystophyllum, the correct name for Neckera Turland N (2013) The Code decoded. A user’s guide to the International Code of Nomenclature for algae, fungi, and plants. Regnum Vegetabile 155. Koeltz Botanical Books, Königstein, v + 169 pp. Turland NJ, Wiersema JH, Barrie FR, Gandhi KN, Gravendyck J, Greuter W, Hawksworth DL, Herendeen PS, Klopper RR, Knapp S, Kusber WH, Li DZ, May TW, Monro AM, Prado J, Price MJ, Smith GF, Zamora Señoret JC (2025) International Code of Nomenclature for algae, fungi, and plants (Madrid Code). Accepted by the Twentieth International Botanical Congress, Madrid, Spain, July 2024. Regnum Vegetabile 162. University of Chicago Press, Chicago, xlvii + 303 pp. https://doi.org/10.7208/chicago/9780226839479.001.0001 Wiersema JH, McNeill J, Barrie FR, Buck WR, Demoulin V, Greuter W, Hawksworth DL, Herendeen PS, Knapp S, Marhold K, Prado J, Prud’homme van Reine WF, Smith GF [Eds] (2015) International Code of Nomenclature for algae, fungi, and plants (Melbourne Code) adopted by the Eighteenth International Botanical Congress Melbourne, Australia, July 2011. Appendices II−VIII. Regnum Vegetabile 157. Koeltz Botanical Books, Königstein, xix + 492 pp. Wijk van der R, Margadant WD, Florschütz PA (1964) Index muscorum. Vol. 3 (Hypnum − O). Regnum Vegetabile 33. International Bureau for Plant Taxonomy and Nomenclature of the International Association for Plant Taxonomy, Utrecht – Netherlands, 529 pp. Wijk van der R, Margadant WD, Florschütz PA (1969) Index muscorum. Vol. 5 (T − Z, Appendix). Regnum Vegetabile 65. International Bureau for Plant Taxonomy and Nomenclature of the International Association for Plant Taxonomy, Utrecht – Netherlands, xii + 922 pp. Wilson KL (2023) Report of the General Committee: 28. Taxon 72(6): 1353–1354. https://doi.org/10.1002/tax.13100 Zijlstra G (1998) Report of the Committee for Bryophyta: 4. Taxon 47(4): 859–862. https://doi.org/10.2307/1224193 Zijlstra G (1999) Report of the Committee for Bryophyta: 5. Taxon 48(4): 815–816. https://doi.org/10.2307/1223655 Zijlstra G (2015) Type species or typical species. Taxon 64(3): 582–585. https://doi. org/10.12705/643.11