Notulae to the Italian native vascular flora: 20

Notulae to the Italian native vascular flora: 20 Fabrizio Bartolucci1, Gianniantonio Domina2, Claudia Angiolini3, Carlo Argenti4, Gianluigi Bacchetta5, Davide Barberis6, Giuseppe Bertotto7, Gianmaria Bonari3,8, Giacomo Calvia9, Francesco Candini3,10, Furio Coltri11, Fabio Conti1, Emanuele Del Guacchio12, Romeo Di Pietro13, Francesco Festi14, Mauro Fois5, Luigi Forte15,16, Gabriele Galasso11, Marcellino Gallo Splendore17, Jacob Koopman18, Michele Lonati19, Francesco Mascia3, Francesco Minutillo20, Juri Nascimbene21, Ginevra Nota19, Giuseppe Pappagallo22, Giuseppe Pellegrino15, Lina Podda5, Giovanni Roffarè23, Alessandro Ruggero5, Federico Selvi24, Giuseppe Silletti25, Adriano Soldano26, Carlo Terranova27, Giancarlo Tondi28, Roberta Vallariello12, Katia Zanatta29, Lorenzo Lastrucci30 1Centro Ricerche Floristiche dell’Appennino (Università di Camerino - Parco Nazionale del Gran Sasso e Monti della Laga), Via Prov.le km 4,2 - San Colombo, 67021 Barisciano (L’Aquila), Italy 2Dipartimento di Scienze Agrarie, Alimentari e Forestali (SAAF), Università di Palermo, Viale delle Scienze, edificio 4, 90128 Palermo, Italy 3Dipartimento di Scienze Ambientali, Università di Siena, Via P.A. Mattioli 4, 53100 Siena, Italy 4Via Pietriboni 7, 32100 Belluno, Italy 5Centro Conservazione Biodiversità (CCB), Dipartimento di Scienze della Vita e dell’Ambiente, Università di Cagliari, viale Sant’Ignazio da Laconi 13, 09123 Cagliari, Italy 6Ente di Gestione delle Aree protette delle Alpi Marittime, Piazza Regina Elena 30, 12010 Valdieri (Cuneo), Italy 7Via Ippolito Nievo 29, 07026 Olbia (Olbia-Tempio), Italy 8National Biodiversity Future Center (NBFC), Piazza Marina 61 900133 Palermo, Italy 9 Facoltà di Scienze Agrarie, Ambientali e Alimentari, Libera Università di Bolzano, Piazza Università 5, 39100 Bolzano, Italy 10Diartimento Scienze della Terra e del Mare, Università di Palermo, Via Archirafi, 22 90123 Palermo, Italy 11Sezione di Botanica, Museo di Storia Naturale di Milano, Corso Venezia 55, 20121 Milano, Italy 12Dipartimento di Biologia, Università di Napoli Federico II, c/o Orto Botanico, Via Foria, 223, 80139 Napoli, Italy 13Dipartimento di Pianificazione, Design, Tecnologia dell’Architettura, Sapienza Università di Roma, Via Flaminia 72, 00196 Roma, Italy 14 Fondazione Museo Civico di Rovereto – Società Museo Civico di Rovereto, Borgo S. Caterina 41, 38068, Rovereto, Trento, Italy 15Dipartimento di Bioscienze, Biotecnologie e Ambiente, Campus Universitario “E. Quagliariello”, Università degli Studi di Bari “Aldo Moro”, Via Orabona 4, 70125 Bari, Italy 16 Museo Orto Botanico, Campus Universitario “E. Quagliariello”, Università degli Studi di Bari “Aldo Moro”, Via Orabona 4, 70125 Bari, Italy 17Museo del Castagno, via Vittorio Emanuele, 87013 Fagnano Castello (Cosenza), Italy 18ul. Kochanowskiego 27, 73-200 Choszczno, Poland 19Dipartimento di Scienze Agrarie, Forestali e Alimentari, Università degli Studi di Torino, Largo Paolo Braccini 2, 10095 Grugliasco (Torino), Italy 20Via Cuostile 5, 04024 Gaeta (Latina), Italy 21BIOME Lab, Dipartimento di Scienze Biologiche, Geologiche e Ambientali, Alma Mater Studiorum - Università di Bologna, Via Irnerio 42, 40126 Bologna, Italy 22Istituto di Ricerca sulle Acque (CNR-IRSA), Via F. De Blasio 5, 70132 Bari, Italy 23Giardino Botanico “Giangio Lorenzoni”, via Cansiglio, 32010 Pian del Cansiglio - Tambre (Belluno), Italy 24Dipartimento di Scienze Agrarie, Alimentari, Ambientali e Forestali, Università di Firenze, Piazzale Italian Botanist 20: 89–107 (2025) doi: 10.3897/italianbotanist.20.179511 https://italianbotanist.pensoft.net Copyright Fabrizio Bartolucci et al.. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited. FLORA AND FUNGA RECORDS Fabrizio Bartolucci et al. / Italian Botanist 20: 89–107 (2025) 90 Cascine 28, 50144 Firenze, Italy 25Via Gioia 80, Santeramo in Colle (Bari), Italy 26Largo Brigata Cagliari, 6, 13100 Vercelli, Italy 27Via G. V. Gravina, 35, 87013 Fagnano Castello (Cosenza), Italy 28Via F. D’Ovidio 89, 00137 Roma, Italy 2931040 Giavera del Montello (Treviso), Italy 30Sistema Museale di Ateneo, Università di Firenze, Via G. La Pira 4, 50121 Firenze, Italy Corresponding author: Fabrizio Bartolucci ([email protected]) Academic editor : Lorenzo Peruzzi|Received 24 November 2025|Accepted 30 November 2025|Published 5 December 2025 Citation: Bartolucci F, Domina G, Angiolini C, Argenti C, Bacchetta G, Barberis D, Bertotto G, Bonari G, Calvia G, Candini F, Coltri F, Conti F, Del Guacchio E, Di Pietro R, Festi F, Fois M, Forte L, Galasso G, Gallo Splendore M, Koopman J, Lonati M, Mascia F, Minutillo F, Nascimbene J, Nota G, Pappagallo G, Pellegrino G, Podda L, Roffarè G, Ruggero A, Selvi F, Silletti G, Soldano A, Terranova C, Tondi G, Vallariello R, Zanatta K, Lastrucci L (2025) Notulae to the Italian native vascular flora: 20. Italian Botanist 20: 89–107. https://doi.org/10.3897/italianbotanist.20.179511 Abstract In this contribution, new data concerning the distribution of native vascular flora in Italy are presented. It includes new records, confirmations, and exclusions to the Italian administrative regions for taxa in the genera Aethusa, Alchemilla, Astragalus, Avena, Carex, Centaurea, Centranthus, Chenopodiastrum, Crepis, Elatine, Filago, Gagea, Lamium, Lappula, Lisymachia, Nymphoides, Onobrychys, Plantago, Pseudopodospermum, Salix, Seseli, Stachys, Ulmus. A new combination in the genus Valeriana (V. pontecorvi) is proposed. Nomenclatural and distribution updates, published elsewhere, and corrigenda are provided as Suppl. material 1. Keywords Endemic taxa, floristic data, Italy, nomenclature, taxonomy How to contribute The text for the new records, exclusions, and confirmations should be submitted electronically to Lorenzo Lastrucci (lorenzo[email protected]). The corresponding specimen along with its scan or photograph have to be sent to FI Herbarium: Sezione di Botanica “Filippo Parlatore” del Museo di Storia Naturale, Via G. La Pira 4, 50121 Firenze (Italy). Those texts concerning nomenclatural novelties and typifications (only for accepted names) should be submitted electronically to: Fabrizio Bartolucci (fabrizio. [email protected]). Each text should be within 1,000 characters (spaces included). Floristic records of native taxa Aethusa cynapium L. subsp. cynapium (Apiaceae) − CAM: Subspecies to be excluded from the flora of Campania. Notulae to the Italian native vascular flora: 20 91 Aethusa cynapium L. subsp. elata (J.Friedl. ex Hoffm.) Schübl. & G.Martens (Apiaceae) ? CAM: Subspecies doubtfully occurring in Campania. 0 LAZ: Subspecies to be confirmed for Lazio. In the framework of the botanical voyages for the Flora napolitana project, Casale and Gussone (1811a) reported “Aethusa cynapium” for the province of Avellino. All the following reports for the same area (Casale 1901, under the name A. cynapium; Fiori 1925, under the name A. cynapium var. domestica Wallr.; Pignatti 1982), very likely also that in Cassitto (1837, under the name A. cynapium), are based on them. The same authors reported A. cynapium also for the cultivated and hilly places of the District of Castellammare (Peninsula of Sorrento) (Casale and Gussone 1811b). Therefore Tenore (1811: XIX) listed A. cynapium without any further detail first for the Kingdom of Naples and later for the province of Naples (Tenore 1823). Later, he became aware that the previous indications were to be referred to A. cynapium subsp. elata instead (Tenore 1824−1829, under the name A. cynapioides). However, he reported this taxon only for northern Abruzzo (based on Gussone 1826). Then, Tenore (1831) did not report anymore A. cynapium for the Kingdom and added to the localities of A. cynapioides also Mt. S. Angelo di Castellammare (Peninsula of Sorrento) (cf. Paoletti 1900; indication not received by Bartolucci et al. 2024). Finally, he recognized the typical A. cynapium in a plant gathered in present-day Lazio (Picinisco, Frosinone) (Tenore 1835−1838). None of the two subspecies has been observed anymore in Campania, and, for example, the indication from the Peninsula was considered as erroneous already by Guadagno (Sched. G780, NAP). In NAP only two specimens are present in Tenore’s collection, both originating from the Botanical Garden of Naples. In the “Gussone Generale” collection, there are no specimens of A. cynapium subsp. cynapium from Italy and none A. cynapium subsp. elata from modern Campania. On the contrary, a sample collected in 1834 in the same place in Lazio cited by Tenore (1835–1838) for A. cynapium, was correctly labelled by Gussone as A. cynapioides (NAP0007046) and so reported in Tenore and Gussone (1843). E. Del Guacchio, R. Vallariello Alchemilla lasenii S.E.Fröhner (Rosaceae) + LAZ: Accumoli (Rieti), Agro Nero - Illica (WGS84: 42.701621°N, 13.309296°E), conca umida torbosa, suolo arenaceo, 1510 m, 31 July 2024, Leg. G. Tondi, R. Di Pietro, Det. F. Festi (FI; Herb. Tondi). – Species new for the flora of Lazio. The finding of this population significantly extends the disjunct Apennine range of this endemic Alpine species (Buccomino et al. 2023). F. Festi, R. Di Pietro, G. Tondi Alchemilla tirolensis Buser (Rosaceae) + LAZ: Accumoli (Rieti), Agro Nero - Illica (WGS84: 42.701621°N, 13.309296°E), conca umida torbosa, suolo arenaceo, 1510 m, 31 July 2024, Leg. G. Tondi, R. Di Fabrizio Bartolucci et al. / Italian Botanist 20: 89–107 (2025) 92 Pietro, Det. F. Festi (FI; Herb. Tondi); ibidem, (WGS84: 42.705304°N, 13.324429°E), prato montano fresco, scendendo dal Lago Secco alla Sorgente dell’Agro Nero, lungo il sentiero 375; suolo arenaceo, 1520 m, 18 June 2023, Leg. G. Tondi, Det. G. Tondi, Conf. F. Festi (Herb. Tondi); Picinisco (Frosinone), Monte Cavallo (WGS84: 41.651163°N, 13.959077°E), prato acquitrinoso, presso la Sorgente del Maresciallo, suolo calcareo, 1700 m, 28 June 2022, Leg. G. Tondi, F. Minutillo, Det. F. Festi (Herb. Tondi). – Species new for the flora of Lazio. The finding of this population extends the range of the species southwards. F. Festi, R. Di Pietro, G. Tondi Astragalus hypoglottis L. subsp. gremlii (Burnat) Greuter & Burdet (Fabaceae) + PUG: Biccari (Foggia), a monte di Lago Pescara (WGS84: 41.3669504°N, 15.1694186°E), Scarpata al margine del sentiero, 985 m, 22 April 2025, Leg. G. Pellegrino, Det. L. Forte, G. Pellegrino (FI, BI No. 60261). – Species and subspecies new for the flora of Puglia. This taxon was previously reported from the Monti Dauni, specifically in the Monte Cornacchia area, although with some doubts, as it was observed exclusively in the fruiting stage (Wagensommer et al. 2014). L. Forte, G. Pellegrino Avena barbata Pott ex Link (Poaceae) + VDA: Montjovet (Aosta), Frazione Bocard, lungo l’autostrada Torino-Aosta, (WGS84: 45.714727°N, 7.670527°E), scarpata autostradale, 381 m, 21 May 2025, M. Lonati, G. Nota (FI). – Species new for the flora of Valle d’Aosta. The species has also been observed in other locations of the lower Aosta Valley, such as in the municipalities of Verrès and Arnad. M. Lonati, G. Nota Carex buxbaumii Wahlenb. (Cyperaceae) − ABR: Species to be excluded from the flora of Abruzzo. The records of this species by Conti et al. (1990, 2019, 2020) should be referred to C. hartmaniorum A.Cajander (see below). Carex hartmaniorum A.Cajander (Cyperaceae) + ABR: Palena (Chieti), Quarto del Barone, prati umidi, 1300 m, 23 May 2000, F. Conti (APP No. 10854); Ovindoli (L’Aquila), Val D’Arano, 1300–1350 m, 23 July 2001, G. Ciaschetti (APP No. 68475); Rivisondoli (L’Aquila), Quarto del Barone, prati umidi, 1250 m, 19 July 2015, G. Ciaschetti (Herb. Maiella); Palena (Chieti), Quarto di S. Chiara, prati umidi, 1250 m, 19 June 2016, G. Ciaschetti (Herb. Maiella, three sheets); Ovindoli (L’Aquila), Val D’Arano (WGS84: 42.130005°N, 13.555773°E), prati umidi, 1330 m, 18 June 2025, F. Conti (APP No. 73375); Ovindoli (L’Aquila), Piana di Ovin- Notulae to the Italian native vascular flora: 20 93 doli (WGS84: 42.140002°N, 13.525366°E), prati umidi, 1350 m, 18 June 2025, F. Conti (APP nos 73377, 73378, 73380; FI). – Species new for the flora of Abruzzo. Thanks to the review of the herbarium material preserved in APP and Herb. Maiella, previously identified as C. buxbaumii (Conti et al. 1990, 2019, 2020), it was possible to ascertain the presence of C. hartmaniorum, new for the Apennines. F. Conti, F. Bartolucci, J. Koopman Centaurea dichroantha A.Kern. (Asteraceae) + VEN: Soverzene (Belluno), Val Gallina (WGS84: 46.21881°N, 12.30323°E), magredo, 425 m, 6 July 2024, Leg. et Det. G. Roffarè, C. Argenti, Conf. F. Martini (FI, Herb. Argenti). – Species confirmed for the flora of Veneto. This species is reported for Veneto in province of Belluno (“segnalata anche in Comelico”, Pignatti 1982; Pignatti et al. 2018). This record was not considered in Argenti et al. (2019) and has never been confirmed. G. Roffarè, C. Argenti Centaurea jacea L. subsp. angustifolia (DC.) Gremli (Asteraceae) + BAS: Policoro (Matera), Bosco Pantano Soprano (WGS84: 40.1718195°N, 16.6487746°E), incolto umido argilloso, 20 m, 27 September 2023, Leg. G. Pellegrino, Det. G. Pellegrino, L. Forte (FI; BI Nos 59478, 59479, 59480). – Subspecies new for the flora of Basilicata. This subspecies was recently reported from Puglia (Bartolucci et al. 2023), and was discovered in Basilicata during fieldwork for a MSc Thesis (Pellegrino 2025). G. Pellegrino, L. Forte Centranthus angustifolius (Mill.) DC. subsp. angustifolius (Valerianaceae) +TOS: Molazzana (Lucca), Alpi Apuane, anello del Monte Rovaio, macereto a ridosso del sentierino verso Trescola, 980 m (WGS84: 44.058888°N, 10.338611°E), 19 June 2024, A. Soldano 20011 (FI). – Species new for the flora of Toscana. This species was not recorded in the recent flora of the Apuan Alps (Pierini and Peruzzi 2025). A. Soldano Chenopodiastrum hybridum (L.) S.Fuentes, Uotila & Borsch +TOS: Sorano (Grosseto), loc. Palazzina (WGS84: 42.711669°N, 11.800137°E), comunità pioniere nitrofile a fenologia tardo primaverile-estiva sviluppate su colture sarchiate non irrigue, 471 m s.l.m., 05 June 2025, Leg. F. Mascia, Det. F. Mascia & C. Angiolini (FI, SIENA). – Species confirmed for the flora of Toscana. This species is reported for Toscana (Bartolucci et al. 2024); however, the related records mostly refer to historical findings, and only two recent records exist for the Fabrizio Bartolucci et al. / Italian Botanist 20: 89–107 (2025) 94 Province of Prato (Stampi 1967; Gestri 2009), one of which is considered doubtful (Peruzzi and Bedini 2013). In both cases, subsequent floristic studies for the same locations did not confirm the presence of the species (Gestri and Peruzzi 2016; Gestri and Lazzeri 2021). No recent herbarium specimens are available at FI herbarium. The present finding represents the unique recent record supported by herbarium specimens confirming the occurrence of the species in the regional territory. F. Mascia, C. Angiolini Crepis neglecta L. subsp. corymbosa (Ten.) Nyman (Asteraceae) + LAZ: Ceprano (Frosinone) (WGS84: 41.552337°N, 13.457593°E), schiarita all’interno del querceto di Via Comuni, suolo arenaceo, 137 m, 11 May 2024, Leg. F. Minutillo, G. Tondi, Det. F. Minutillo (FI; Herb. Tondi). – Subspecies new for the flora of Lazio. F. Minutillo, G. Tondi Elatine brochonii Clavaud (Elatinaceae) + ITALIA (SAR): La Maddalena (Sassari), Caprera, Fosso Stefano (WGS84: 41.219087°N, 9,464109°E), rive sabbiose di bacino artificiale, Leg. A. Ruggero, A. Di Giacomo, Det. A. Ruggero, G. Bacchetta (CAG). – Species new for the Italian flora (Sardegna). This species is native to the Mediterranean-Atlantic area, and is characterized by small and limited populations in Algeria (de Bélair 2005; Rhazi et al. 2007), Corsica (Lorenzoni and Paradis 1997; Delage and Hugot 2020), continental France (Chammard et al. 2019), Morocco (Rhazi et al. 2007), Portugal (Porto et al. 2012) and Spain (Cirujano and Velayos 1996; Benito Alonso 2010; Lopez Udias and Fabregat Llueca 2011). The only population found in Sardegna grows on the banks of an artificial reservoir and appears in late spring, when the freshwater surface area shrinks, leaving large portions free for the development of hygrophilous and heliophilous therophytic vegetation belonging to the Isoëtion Br.-Bl. 1936 alliance, and more specifically to the Elatino brochonii-Juncetum pygmaei Lorenzoni & Paradis ex Benito Alonso 2010 association (Reymann et al. 2016). Elatine brochonii colonises wet sands just above the water level, in a small corner of the reservoir. G. Bacchetta, A. Ruggero Filago congesta Guss. ex DC. (Asteraceae) + SAR: Cagliari, promontorio di Cala Mosca, sul sentiero per la Sella del Diavolo (WGS84: 39.183661°N, 9.157292°E), 50–80 m s.l.m., 30 March 2025, G. Calvia (FI, Herbarium Calvia). – Species confirmed for the flora of Sardegna. This species was reported in Sardegna by Fiori (1927), who referred to a herbarium specimen collected by Marcucci in 1866 on the hill of Bonaria (Cagliari), preserved in FI. Later, Arrigoni (2015) considered it as possibly extinct on the island, and Bartolucci et al. (2018, 2024) did not confirm its presence. However, this species still persists on the calcareous hills of Cagliari, particularly in areas such as Monte Urpinu and Capo Sant’Elia, in barren and open habitats, where it sometimes grows alongside F. pygmaea. G. Calvia, F. Mascia Notulae to the Italian native vascular flora: 20 95 Gagea minima (L.) Ker Gawl. (Liliaceae) + ABR: Rocca di Mezzo (L’Aquila), V. di Fùndoli (WGS84: 42.184331°N, 13.557976°E), radure e margine faggeta, 1378 m, 2 May 2025, F. Bartolucci (APP No. 73371; FI). – Species new for the flora of Abruzzo. F. Bartolucci, F. Conti Lamium bifidum Cirillo subsp. bifidum (Lamiaceae) + VEN: Villorba (Treviso) (WGS84: 45.74955°N, 12.21524°E), nitrophilous margin of a hedge, mainly composed of Sambucus nigra and Robinia pseudoacacia, along a road; the site is in the high venetian plain area characterized by low naturalness, high population density and intensive agriculture, 54 m a.s.l., 23 March 2025, Leg. K. Zanatta, Det. K. Zanatta, J. Nascimbene (FI; Herb. Zanatta). – Subspecies new for the flora of Veneto. This species has a Steno-Mediterranean distribution and an almost ruderal ecology. Both climate warming and human disturbance could have favored its northward migration in Italy. K. Zanatta, J. Nascimbene Lappula squarrosa (Retz.) Dumort. (Boraginaceae) + TOS: Firenze (Firenze), Via Romana all’incrocio con Via de’ Mori (WGS84: 43.763193°N, 11.244352°E), spigolo tra il muro e la strada, 90 m, 8 May 2025, F. Selvi (FI). – Species confirmed for the flora of Toscana. In the locality here reported, a few plants of this species were growing on the sidewalk close to a wall of Giardino Corsi. The only known collection of this species from Toscana dates back to 1876 (29 July 1876, A. Mori, MOD) and comes from “Boscolungo” a locality near Abetone in the northern Apennines (province of Pistoia). In the 19th century, this species was sporadically observed in some other localities of the Tuscan Apennines (Lunigiana, Garfagnana), around Lucca and Firenze and possibly upper Maremma, but it was considered possibly extinct in the region (Selvi and Cecchi in Arrigoni 2019) and hence not confirmed by Bartolucci et al. (2024). The present finding is thus a confirmation of the species for the Tuscan flora, though the presence in the historical center of Firenze is most likely casual, of uncertain origin and not stable. F. Coltri, F. Selvi Lisymachia monellii (L.) U.Manns & Anderb. (Primulaceae) − SAR: Species to be excluded from the flora of Sardegna. This species has been reported in Sardegna since Moris (1827), being widely present in the Sassarese and, in particular, in the Iglesiente and on the island of S. Pietro. Both redflowered and blue-flowered specimens have been reported under this epithet, as this trait was considered part of intraspecific variation. However, molecular analyses by Jiménez-López et al. (2022) have demonstrated that the two-colour typologies constitute independent Fabrizio Bartolucci et al. / Italian Botanist 20: 89–107 (2025) 96 species: L. monellii (blue flowers) and L. collina (Schousb.) F.J.Jiménez López (red flowers). Only red-flowered individuals are present in Sardegna, which therefore refer to L. collina. M. Fois, L. Podda Plantago altissima L. (Plantaginaceae) + UMB: Foligno (Perugia), Palude di Colfiorito, Parco Regionale di Colfiorito (WGS84: 43.018341°N, 12.871559°E), prato umido periodicamente allagato a margine della palude, 755 m, 30 April 2025, Leg. G. Bonari, F. Candini, V. GonzálezGarcía, Det. G. Bonari, V. González-García (FI). – Species new for the flora of Umbria. G. Bonari, F. Candini Pseudopodospermum hispanicum (L.) Zaika, Sukhor. & N.Kilian subsp. neapolitanum (Grande) Bartolucci, Galasso & F.Conti (Asteraceae) + PUG: Gravina in Puglia (Bari), Bosco Selva (WGS84: 40.756224°N, 16.492808°E), margine boschivo, 438 m, 17 May 2025, G. Silletti, det. F. Bartolucci, G. Silletti (APP Nos 73381, 73383; FI). – Subspecies confirmed for the flora of Puglia. This subspecies was formerly indicated by Trotter and Romano (1914, under the name Scorzonera hispanica L. var. trachisperma) for “Bosco Aquara” at Monte Crispigliano (Daunia Mountains). F. Bartolucci, G. Silletti Salix eleagnos Scop. (Salicaceae) + PUG: Orsara di Puglia (Foggia), località Le Querce/C. Forestale (WGS84: 41.265599°N, 15.339151°E), lungo il corso superiore del torrente Cervaro (Monti Dauni) in area inondabile immediatamente all’esterno della fascia ripariale arborata, altitudine 380 m, esposizione Sud, 21 June 2025, G. Pappagallo (FI). – Species new for the flora of Puglia. G. Pappagallo Seseli annuum L. subsp. annuum (Apiaceae) − ABR: Species to be excluded from the flora of Abruzzo. The record of this species (Conti et al. 2025) should be referred to S. pallasii Besser (see below). Seseli pallasii Besser (Apiaceae) + ABR: Terranera (L’Aquila), nei pressi dell’abitato lungo la SP38 (WGS84: 42.237641°N, 13.52913°E), pascoli aridi pietrosi, 1292 m, 23 September 2024, F. Bartolucci (APP No. 72771); ibidem, 12 Jul 2025, F. Bartolucci (APP No. 73467; FI). – Species new for the flora of Abruzzo. F. Bartolucci, F. Conti Notulae to the Italian native vascular flora: 20 97 Stachys romana (L.) E.H.L.Krause (Lamiaceae) + PIE: Alto (Cuneo), sotto Strada Provinciale 216 (WGS84: 44.108629°N, 7.995099°E), prateria xerica, 760 m, 8 April 2025, Leg. D. Barberis, Det. D. Barberis, G. Marengo, G. Nota (FI). – Species confirmed for the flora of Piemonte. This species was reported in Piemonte as an erroneous record (Bartolucci et al. 2024). The present finding at the described locality, along with another site in the municipality of Alto (WGS84: 44.108629°N, 7.995099°E) is near the Ligurian border. D. Barberis, G. Nota Ulmus minor Mill. subsp. canescens Bartolucci & Galasso (Ulmaceae) + BAS: Policoro (Matera), Bosco Pantano Soprano (WGS84: 40.1711897°N, 16.6489122°E), bosco misto planiziale, 20 m, 30 September 2024, Leg. G. Pellegrino, Det. G. Pellegrino, L. Forte (BI No. 59518, FI). – Subspecies new for the flora of Basilicata. This subspecies was detected during fieldwork for a MSc Thesis (Pellegrino 2025). G. Pellegrino, L. Forte Floristic records of regional alien taxa Crepis foetida L. subsp. rhoeadifolia (Asteraceae) + (NAT) LAZ: Lago di Posta Fibreno (Posta Fibreno, Frosinone) (WGS84: 41.694300°N, 13.692528°E), incolto al bordo meridionale del lago; suolo calcareo, 290 m, 2 July 2022, Leg. F. Minutillo, G. Tondi, Det. G. Galasso (FI; Herb. Tondi). – Naturalized regional alien subspecies new for the flora of Lazio. The species was recently recorded as casual alien by Pinzani et al. (2025). F. Minutillo, G. Tondi Nymphoides peltata (S.G.Gmel.) Kuntze (Menyanthaceae) + (NAT) CAL: Fagnano Castello (Cosenza), lago della Paglia (WGS84: 39.560946°N, 16.018152°E), presso la sponda settentrionale del bacino, 1.061 m s.l.m., 14 giugno 2025, C. Terranova, M. Gallo Splendore (FI). – Naturalized regional alien species new for the flora of Calabria. Nymphoides peltata is distributed in central-northern Italy and Sardegna (Bartolucci et al. 2024). This species was first detected in Calabria four years ago with a single individual occupying an area with a diameter of approximately 1 m2. To date, the species is expanding and the area it occupies exceeds 50 square metres. The geographic remoteness of other Italian populations could exclude the spread through hydrochory and ectozoochory (Cook 1990) and intentional introduction by humans is plausible. C. Terranova, M. Gallo Splendore Fabrizio Bartolucci et al. / Italian Botanist 20: 89–107 (2025) 104 Gestri G, Pierini B, Peruzzi L, Festi F (2022) Aggiornamento sulla distribuzione del genere Alchemilla L. (Rosaceae) in Toscana (Italia centrale). Atti della Società Toscana di Scienze Naturali di Pisa, Memorie Serie B 129: 47–51. Gestri G, Pierini B, D’Antraccoli M, Bernardini A, Peruzzi L (2023) An updated inventory of the vascular flora of the Cerbaie hills (Tuscany, Italy). Italian Botanist 15: 165–175. https://doi.org/10.3897/italianbotanist.15.105302 Grillini P, Centurione N (2023) Le orchidee della Valle del Sillaro (Asparagales: Orchidaceae). Quaderno di Studi e Notizie di Storia Naturale della Romagna 58: 153–164. Guarino R, Vynokurov D, Dengler J, Berastegi A, Biurrun I, Garcia-Mijangos I, Janišová M, Lonati M, Nikolei R, Mainetti A, Miskova O, Sánchez-Carmona S, Tyshchenko O, Nota G (2024) (Re)discovery of four annual vascular plant species in dry grasslands in the Southwestern Alps (NW Italy). Palaearctic Grasslands 62: 28–33. https://doi.org/10.21570/ EDGG.PG.62.28-33 Gussone G (1826) Plantae rariores quas in itinere per oras Jonii ac Adriatici Maris ert per regionises Samnii et Aprutii collegit Joannes Gussone. Ex Regia Typographia, Neapoli [Naples]. https://doi.org/10.5962/bhl.title.44889 Iamonico (2025) Clarifications about the identity of the names Sagina alexandrae and S. hawaiensis (Caryophyllaceae). Comprehensive Plant Biology 49(2): 375–379. https://doi. org/10.2298/CPB2502375I Iamonico D, Verloove F, De Mei S (2025) Uncovering Cyperus polystachyos in Europe: nomenclatural insights and new historical records. Plants 14: 3270. https://doi.org/10.3390/ plants14213270 Jiménez-López FJ, Viruel J, Arista M, Ortiz PL, Talavera M (2022) Molecular approaches reveal speciation between red and blue flowered plants in the Mediterranean Lysimachia arvensis and Lysimachia monelli (Primulaceae). Botanical Journal of the Linnean Society 199(2): 557–577. https://doi.org/10.1093/botlinnean/boab081 Kadereit JW (2025) A reversed subspecific classification of Moehringia bavarica (Caryophyllaceae). Willdenowia 55: 95–105. https://doi.org/10.3372/wi.55.09 Kozlowski G, Freigang J, Pasta S, Guarino R, Anania F, Champoud L, Garfì G, Silvestre Gristina A, Marcenò C, Picone RM, Crisafulli A (2025) Update on the current occurrence of two rare fern species in Sicily (Italy): Dryopteris affinis and D. robusta in the Peloritani Mountains. Mediterranean Botany 46(2): e99974. https://doi.org/10.5209/mbot.99974 Leoni G, Perico G (2025) 174. Scorzoneroides montana (Lam.) Holub subsp. melanotricha (Vierh.) Gutermann (Asteraceae) – Dente di leone montano. In: Perico G (Ed.) Segnalazioni floristiche. XXIV contributo (167÷186). Notiziario Floristico del Gruppo Flora Alpina Bergamasca 67: 12. López Udias S, Fabregat Llueca C (2011) Nuevos datos para la flora de Aragón. Flora Montiberica 49: 85–95. Lorenzoni C, Paradis G (1997) Description phytosociologique d’une mare temporaire à Elatine brochonii dans le Sud de la Corse. Bulletin de la Société Botanique du Centre-Ouest N.S. 28: 21–46. Lyskov D, Akalın E, Bagheri A, Terentieva E, Tuncay HO, Kroupsky I, Samigullin T (2025) Break it and better it: molecular and morphological data support the split of Ferulago (Api- Notulae to the Italian native vascular flora: 20 105 aceae) into four genera. Botanical Journal of the Linnean Society: boaf049. https://doi. org/10.1093/botlinnean/boaf049 Moris GG (1827) Stirpium sardoarum elenchus. Ex Typiis Regiis, Carali. Mosyakin SL, Fedoronchuk MM, McNeill J (2025) Simplifying the nomenclature of Sorbus sensu lato: new nomenclatural solutions in Aria and Hedlundia (Rosaceae). Ukrainian Botanical Journal 82(3): 206–224. https://doi.org/10.15407/ukrbotj82.03.206 Ottavi GA (1862) Sui mezzi di migliorare l’agricoltura delle regioni meridionali ed in specie di quella di Sardegna. Tipografia Giuseppe Nani, Casale. Pellegrino G (2025) Indagine floristica del Bosco Pantano Soprano di Policoro (Basilicata). M.S. Thesis, Natural and Environmental Science, Università degli Studi di Bari “Aldo Moro”, Italy. Peruzzi L, Bedini G [Eds] (2013) Wikiplantbase #Toscana. http://bot.biologia.unipi.it/wpb/ toscana/index.html Peruzzi L, Pierini B, Arduini I, Bedini G, Franzoni J (2025) The vascular flora of Pisa (Tuscany, Central Italy). Plants 14(3): 307. https://doi.org/10.3390/plants14030307 Paoletti G (1900) Umbelliferae. In: Fiori A, Paoletti G (Eds) Flora analitica d’Italia, Vol. 2(1): 138–210. Tipografia del Seminario, Padova [Padua]. Pierini B, Peruzzi L (2025) The vascular flora of the Apuan Alps (Tuscany, Central Italy). Italian Botanist 19: 45–63. https://doi.org/10.3897/italianbotanist.19.145947 Pignatti S (1982) Flora d’Italia, 1–3. Edagricole, Bologna. Pignatti S, Guarino R, La Rosa M [Eds] (2017) Flora d’Italia, Ed. 2, Vol. 1. Edagricole, Milano. Pignatti S, Guarino R, La Rosa M [Eds] (2018) Flora d’Italia, Ed. 2, Vol. 3. Edagricole, Milano. Pinzani L, Di Lernia D, Pelella E, Ceschin S (2025) The vascular flora of Italian volcanic lake calderas: a comprehensive floristic study. Environments 12(9): 327. https://doi. org/10.3390/environments12090327 Porto M, Pereira AJ, Rego R, Farminhão J, Clemente A, Vitorino JL, Tauleigne-Gomes C (2012) Elatine brochonii Clavaud (Elatinaceae), a new species to the Portuguese flora. Acta Botanica Malacitana 37: 173–178. https://doi.org/10.24310/abm.v37i0.2688 POWO (2025) [onwards] Plants of the World Online. Facilitated by the Royal Botanic Gardens, Kew. https://powo.science.kew.org/ [Accessed: 19.10.2025] Ravasio G (2025) 181. Anacamptis berica Doro (Orchidaceae) – Orchide dei Colli Berici. In: Perico G (Ed.) Segnalazioni floristiche. XXIV contributo (167÷186). Notiziario Floristico del Gruppo Flora Alpina Bergamasca 67: 14. Reymann J, Panaiotis C, Bioret F, Bacchetta G, Delage A, Delbosc P, Gamisans J, Gauberville C, Hugot L, O’Deye-Guizen K, Piazza C, Pioli A (2016) Prodrome des végétations de Corse. Document Phytosociologiques, serie 3, 4: 1–175. Rhazi L, Grillas P, Charpentier A, Rhazi M, Leclainche N, Titolet D, Desnoubes L, Duborper E, Yavercovski N, El Khyari D (2007) Effet de l’hydrologie sur les populations d’une espèce rare des mares temporaires méditerranéennes: Elatine brochonii (Elatinaceae). In: Paracuellos M (Ed.), Ambientes Mediterraneos. Funcionamiento, biodiversidad y conservación de los ambientes mediterráneos. Colección Medio Ambiente 2: 185–193. [Almeria] Scafidi F (2025) The genus Adenostyles Cass. (Asteraceae) in Southern Italy and Sicily. Biodiversity Journal 16(3): 415–422. https://doi.org/10.31396/Biodiv.Jour.2025.16.3.415.422 Fabrizio Bartolucci et al. / Italian Botanist 20: 89–107 (2025) 106 Šegota V, Gligora Udovič M, Levkov Z, Bogdanović S, Rimac A, Alegro A, Šušnjara M, Doboš M, Temunović M, Žutinić P, Budinski I, Conti F, Dorić V, Kulaš A, Bučar M, Cindrić M, Engelen A, Ilić B, Jeričević M, Maslać Mikulec M, Šabanović E, Vuković N, BorovečkiVoska Lj, Ćato S, Ćetković I, Hudina T, Huška D, Jantol N, Jeričević N, Limić I, Loos U, Klarić M, Kovačević M, Rogošić M, Škunca L, Tkalčec Z, Jurina D, Stenger-Kovács C (2025) New findings of rare and under-recorded vascular plants, bryophytes, lichens, fungi and algae in Croatia and neighbouring countries – 2. Journal of the Croatian Botanical Society 13(1): 72–109. https://doi.org/10.46232/glashbod.13.1.5 Selvaggi A, Soldano A, Pascale M, Dellavedova R [Eds] (2006) Note floristiche piemontesi n. 13–47. Rivista Piemontese di Storia Naturale 27: 429–441. Selvaggi A, Soldano A, Pascale M, Dellavedova R [Eds] (2025) Note floristiche piemontesi n. 1315–1354. Rivista Piemontese di Storia Naturale 46: 27–47. Selvi F, Martellos S, Conti M (2025) Thirty-five years of floristic collections in southern Tuscany (Italy). Biodiversity Data Journal 13: e160516. https://doi.org/10.3897/BDJ.13.e160516 Sennikov AN, Tikhomirov VN (2024) Atlas Florae Europaeae notes, 35. Further critical notes on Cytisus sect. Tubocytisus (Fabaceae) in Europe. PhytoKeys 238: 199–230. https://doi. org/10.3897/phytokeys.238.118032 Sochor M, Lepší M, Lepší P, Velebil J, Király G, Trávníček B (2025) Taxonomy and nomenclature of Rubus ser. Glandulosi (Rosaceae) across its Eurasian range: Revised concepts and new approaches. Taxon 74: 1106–1152. https://doi.org/10.1002/tax.13380 Spadaro V, Marino P, Scuderi L, Venturella G, Raimondo FM (2024) Biodiversity in some populations of Crataegus (Rosaceae) from western Sicily: Description of two new species and notes on conservation and valorisation. Flora Mediterranea 34: 239–255. https://doi. org/10.7320/FlMedit34.239 Stampi P (1967) Florula infestante e cicli d’infestazione nelle risaie di Poggio a Caiano. Istituto di Botanica, Agraria e Forestale, Università di Firenze. Tenore M (1811) Flora napolitana, Vol. 1(2). Nella Stamperia Reale, Napoli [Naples]. Tenore (1823) Flora medica universale e flora particolare della provincia di Napoli. In: Tenore M, Corso delle Botaniche Lezioni, Vol. 4(1). Tipografia del Giornale Enciclopedico, Napoli [Naples]. Tenore M (1824–1829) Flora napolitana, Vol. 3. Nella Stamperia Francese, Napoli [Naples]. Tenore M (1831) Sylloge plantarum vascularium florae neapolitanae hucusque detectarum. Tipografia del Fibreno, Neapoli [Naples]. Tenore M (1835–1838) Flora napolitana, Vol. 5. Dalla Stamperia e Cartiera del Fibreno, Napoli [Naples]. Tenore M, Gussone G (1843) Rapporto intorno alle peregrinazioni de’ soci ordinari signor M. Tenore e G. Gussone eseguite in Luglio 1834. Atti della Reale Accademia delle Scienze (Napoli) 5(1): 283−334. Tison J-M, de Foucault B (2014) Flora Gallica. Flore de France. Biotope Éditons, Mèze. Trotter A, Romano M (1914) Sulla Flora di M. Crispiniano in Puglia. Nuovo Giornale Botanico Italiano, nuova serie, 21(4): 398–435. Valdés B, Scholz H (2009) [onwards] Poaceae. Euro+Med Plantbase - the information resource for Euro-Mediterranean plant diversity. https://europlusmed.org/ [Accessed: 10.11.2025]. Notulae to the Italian native vascular flora: 20 107 Viciani D, Ferretti G, Bernardini A, Raffaelli M, Lastrucci L (2020) An annotated checklist of the vascular flora of the Arezzo administrative province (Tuscany, Italy). Plant Biosystems 155(5): 971–982. https://doi.org/10.1080/11263504.2020.1810811 Voisin C, Choler P, Vacher C, Perrier C, Renaud J, Rioux D, Boucher FC (2025) Deciphering the drivers of speciation in Alpine Noccaea (Brassicaceae) reveal an evolution predominantly shaped by geography. Molecular Phylogenetics and Evolution 211: 108381. https:// doi.org/10.1016/j.ympev.2025.108381 Wagensommer RP, Marrese M, Perrino EV, Bartolucci F, Cancellieri L, Carruggio F, Conti F, Di Pietro R, Fortini P, Galasso G, Lattanzi E, Lavezzo P, Longo D, Peccenini S, Rosati L, Russo G, Salerno G, Scoppola A, Soldano A, Stinca A, Tilia A, Turco A, Medagli P, Forte L (2014) Contributo alla conoscenza floristica della Puglia: resoconto dell’escursione del Gruppo di Floristica (S.B.I.) nel 2011 nel settore meridionale dei Monti della Daunia. Informatore Botanico Italiano 46(2): 175–208. Supplementary material 1 Additional information Authors: Fabrizio Bartolucci, Gabriele Galasso Data type: pdf Explanation note: 1. Nomenclatural updates; 2. Distribution updates; 3. Synonyms, misapplied or included names. Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited. Link: https://doi.org/10.3897/italianbotanist.20.179511.suppl1