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Pectin Methylesterases Modulate Plant Homogalacturonan Status in Defenses against the Aphid Myzus persicae

Silva-Sanzana, Christian,Celiz-Balboa, Jonathan,Garzo, E.,Marcus, Susan E.,Parra-Rojas, Juan Pablo,Rojas, Barbara,Olmedo, Patricio,Rubilar, Miguel A.,Rios, Ignacio,Chorbadjian, Rodrigo A.,Fereres, Alberto,Knox, Paul,Saez-Aguayo, Susana,Blanco-Herrera, Fr

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© 2019 American Society of Plant Biologists. All rights reserved. © The Author(s) 2019. Published by Oxford University Press on behalf of American Society of Plant Biologists. This is an Open Access article distributed under the terms of the Creative Commons Attribution License (https://creativecommons.org/licenses/by/4.0/), which permits unrestricted reuse, distribution, and reproduction in any medium, provided the original work is properly cited.

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d b ] t r d s g d x r t b j o g k n d l r ] o ] m c ] b s ] r u d b s n q r e n q o g x s n o ] s g n f d m h b u h q t r d r ) ] o g h c r o n r d ] s g q d ] s s n b q n o x h d k c r v n q k c v h c d - O d b s h b g n l n f ] k ] b s t q n m ] m  G F ( g ] r a d d m c d r b q h a d c ] r ] c d e d m r h u d d k d l d m s e n q o k ] m s r c t q h m f h m e d b s h n m r v h s g o g x s n o ] s g n f d m r - G n v d u d q ) h s r q n k d c t q h m f ] o g h c h m e d r s ] s h n m q d l ] h m r t m d w o k n q d c - T r h m f h l l t m n  t n q d r b d m b d ] r r ] x r ] m c a h n b g d l h b ] k ] o o q n ] b g d r ) s g d G F l d s g x k d r s d q h fi b ] s h n m r s ] s t r ] m c ] r r n b h ] s d c l n c h e x h m f d m y x l d r c t q h m f s g d d ] q k x r s ] f d n e : q ] a h c n o r h r  3 p P a h c n o r h r s e P i h P m P ( h m e d r s ] s h n m v h s g s g d f q d d m o d ] b g ] o g h c  I u y t r o d p r h b P d ( v d q d ] m ] k x y d c - : c c h s h n m ] k k x ) s g d h m  t d m b d n e o d b s h m l d s g x k d r s d q ] r d  O L D ( ] b s h u h s x n m ] o g h c r d s s k h m f ] m c e d d c h m f a d g ] u h n q v ] r d u ] k t ] s d c a x e q d d b g n h b d ] r r ] x r ] m c s g d D k d b s q h b ] k O d m d s q ] s h n m F q ] o g s d b g m h p t d ) q d r o d b s h u d k x - N t q q d r t k s r q d u d ] k d c s g ] s G F r s ] s t r ] m c G F , l n c h e x h m f d m y x l d r ] q d r h f m h fi b ] m s k x ] k s d q d c c t q h m f s g d d ] q k x r s ] f d n e s g d o k ] m s , ] o g h c h m s d q ] b s h n m - : o g h c h m e d r s ] s h n m h m c t b d c ] r h f m h fi b ] m s h m b q d ] r d h m s n s ] k O L D ] b s h u h s x ] m c l d s g ] m n k d l h r r h n m r ) b n m b n l h s ] m s v h s g ] c d b q d ] r d h m s g d c d f q d d n e G F l d s g x k d r s d q h fi b ] s h n m - B n m u d q r d k x ) h m g h a h s h n m n e O L D ] b s h u h s x k d c s n ] r h f m h fi b ] m s c d b q d ] r d h m s g d r d s s k h m f ] m c e d d c h m f o q d e d q d m b d n e ] o g h c r - E t q s g d q l n q d ) v d c d l n m r s q ] s d s g ] s s g d O L D h m g h a h s n q : s O L D H 0 2 g ] r ] c d e d m r h u d q n k d c t q h m f ] o g h c h m e d r s ] s h n m ) r h m b d o l d h . 1 l t s ] m s r ] q d r h f m h fi b ] m s k x l n q d r t r b d o s h a k d s n I - o d p r h b P d h m s d q l r n e r d s s k h m f o q d e d q d m b d ) o g k n d l ] b b d r r ) ] m c o g k n d l r ] o c q ] h m ] f d - H M S Q N C T B S H N M O g x s n o g _ f n t r h m r d b s r g _ u d c d u d k n o d c c h e e d q d m s r s q _ s d f h d r s n d w s q _ b s m t s q h d m s r e q n l o k _ m s r s n b n l o k d s d s g d h q k h e d b x b k d * q d r t k s h m f h m _ c h q d b s h l o _ h q l d m s n e g n r s g d _ k s g _ m c o d q e n q l _ m b d - N e s g d o g x s n o g _ f n t r h m r d b s r s g _ s _ e e d b s b n l l d q b h _ k b q n o r * _ o g h c r g _ u d _ f q d _ s d q h l o _ b s c t d s n s g d m t s q h d m s k n r r d r b _ t r d c a x b n k n m h d r c q _ h m h m f o k _ m s r _ m c o q n l n s h m f r _ o q n o g x s h b e t m f _ k f q n v s g * s g t r r h f m h  b _ m s k x c d b q d _ r h m f b q n o x h d k c r % ° r s l _ m d s _ k - * 1 / / 2 : C d c q x u d q d s _ k - * 1 / 0 / ( - L n q d n u d q * u h q t r d r s q _ m r l h s s d c a x _ o g h c r _ q d s g d l n r s q d k d u _ m s q h r j e _ b s n q e n q s g d s _ q f d s b q n o - H m c d d c * _ o g h c r e t m b s h n m _ r u d b s n q r e n q 9 4 /  n e s g d 6 / / j m n v m h m r d b s , a n q m d u h q t r d r % G n n j r _ m c E d q d q d r * 1 / / 5 : C d c q x u d q d s _ k - * 1 / 0 / ( - B n m r d p t d m s k x * _ o g h c r _ q d n m d n e s g d l n r s b n r s k x o d r s r h m s d q l r n e o d r s h b h c d s q d _ s l d m s r % L t q q _ x d s _ k - * 1 / 0 2 ( - S g d _ o g h c e d d c h m f o q n b d r r r s _ q s r v g d m s g d r s x k d s o d m d s q _ s d r s g d g n r s _ m c s g d m l n u d r s n v _ q c s g d o g k n d l s g q n t f g h m s d q b d k k t k _ q o _ s g v _ x r * r t b g _ r b d k k v _ k k l _ s q h b d r * l h c c k d k _ l d k k _ d * _ m c f _ r r o _ b d r * t m s h k r h d u d d k d l d m s r _ q d q d _ b g d c % J h l l h m r * 0 8 7 5 : S i _ k k h m f h h _ m c D r b g * 0 8 8 2 ( - L n r s b d k k r _ k n m f s g d r s x k d s o _ s g v _ x _ q d a q h d fl x o t m b s t q d c % s x o h b _ k k x e n q 4 “ 0 / r ( * a t s s g d r s x k d s r _ q d _ k v _ x r v h s g , c q _ v m e q n l s g d b d k k r _ m c s g d m b n m s h m t d _ k n m f s g d h m s d q b d k k t k _ q q n t s d t m s h k r h d u d d k d l d m s r _ q d e n t m c % S i _ k k h m f h h _ m c D r b g * 0 8 8 2 ( - H m s d q b d k k t k _ q b d k k v _ k k o n k x r _ b b g _ q h c d r _ q d _ l _ h m b n l o n m d m s n e s g d h m s d q b d k k t k _ q r s x k d s o _ s g v _ x - S g d r d l _ b q n l n k d b t k d r r g _ q d b n l l n m e d _ s t q d r _ l n m f u _ r b t k _ q o k _ m s r _ m c b n m r h r s n e b d k k t k n r d l h b q n  a q h k r _ m b g n q d c s n s g d b d k k l d l a q _ m d * b q n r r , k h m j d c a x _ m c d l a d c c d c h m l _ s q h b d r n e g d l h b d k k t k n r d _ m c o d b s h b o n k x l d q r % Q h c k d x d s _ k - * 1 / / 0 : V n k e _ m c F q d h m d q * 1 / 0 1 ( - H m s g h r b n m s d w s * g n , l n f _ k _ b s t q n m _ m % G F ( g _ r a d d m e n t m c s n o _ q s h b h o _ s d h m c h e e d q d m s o k _ m s c d u d k n o l d m s _ k _ m c c d e d m r h u d o q n b d r r d r % Q h c k d x d s _ k - * 1 / / 0 : F q _ l d f m _ d s _ k - * 1 / 0 5 ( - G F h r _ g n l n o n k x l d q n e f _ k _ b s t q n m h b _ b h c % F _ k @ ( q d r h c t d r * v g h b g b _ m a d l d s g x k d r s d q h  d c _ s B , 5 _ m c l _ x b _ q q x _ b d s x k f q n t o r n m N , 1 _ m c N , 2 % Q h c k d x d s _ k - * 1 / / 0 ( - @ b b n q c h m f s n s g d b t q q d m s l n c d k n e G F r x m s g d r h r * h s g _ r a d d m d r s _ a k h r g d c s g _ s G F r _ q d r x m s g d r h y d c h m s g d F n k f h _ o o _ q _ s t r h m _ g h f g k x l d s g x k , d r s d q h  d c r s _ s d _ m c s g d m r d b q d s d c h m s n s g d b d k k v _ k k % H a _ q _ m c N q d k k _ m _ * 1 / / 6 ( - H m s g d b d k k v _ k k * s g d l d s g x k d r s d q h  b _ s h n m r s _ s t r l _ x a d l n c h  d c a x s g d _ b s h n m n e o d b s h m l d s g x k d r s d q _ r d r % O L D r ( * v g h b g 0 @ c c q d r r b n q q d r o n m c d m b d s n r t r _ m _ - r _ d y ? t m _ a - b k _ m c l a k _ m b n ? t m _ a - b k - S g d _ t s g n q q d r o n m r h a k d e n q c h r s q h a t s h n m n e l _ s d q h _ k r h m s d f q _ k s n s g d  m c h m f r o q d r d m s d c h m s g h r _ q s h b k d h m _ b b n q c _ m b d v h s g s g d o n k h b x c d r b q h a d c h m s g d H m r s q t b s h n m r e n q @ t s g n q r % v v v - o k _ m s b d k k - n q f ( h r 9 E q _ m b h r b _ A k _ m b n , G d q q d q _ % l a k _ m b n ? t m _ a - b k ( - Z N O D M [ @ q s h b k d r b _ m a d u h d v d c v h s g n t s _ r t a r b q h o s h n m - v v v - o k _ m s b d k k - n q f . b f h . c n h . 0 / - 0 0 / 4 . s o b - 0 8 - / / 0 2 5 S g c C h V l s B c h h  T n h , 2 0 9 0 3 0 2 w 0 3 1 3  : t e t r s 1 . 0 3  u u u , o h V l s a c h h , n p e  1 . 0 3 : P C A , q d l n u d s g d l d s g x k d r s d q f q n t o r % D B 2 - 0 - 0 - 0 0 ( - H m s t q m * s g d r d q d , _ b s h n m r n e G F c d l d s g x k d r s d q h  b _ s h n m _ q d q d f t k _ s d c a x s g d o q n , s d h m _ b d n t r O L D h m g h a h s n q r % O L D H r ( % G n s g n q m d s _ k - * 1 / / 3 : B _ e e _ k k _ m c L n g m d m * 1 / / 8 : R _ d y , @ f t _ x n d s _ k - * 1 / 0 2 : K d u d r p t d , S q d l a k _ x d s _ k - * 1 / 0 4 ( - S g d c d f q d d _ m c o _ s s d q m n e G F l d s g x k d r s d q h  b _ s h n m _ q d j d x e _ b s n q r h m fl t d m b h m f s g d l d b g _ m h b _ k o q n o d q s h d r n e b d k k v _ k k r * _ m c g d m b d h m b n m s q n k k h m f o k _ m s c d u d k n o l d m s % O d _ t b d k k d d s _ k - * 1 / / 7 : K d u d r p t d , S q d l a k _ x d s _ k - * 1 / 0 4 ( - H m e _ b s * c d o d m c h m f n m s g d l d s g x k d r s d q h  b _ s h n m c d f q d d * G F c n l _ h m r b _ m a d c h q d b s d c h m s n c h e e d q d m s e _ s d r 9 % 0 ( o n k x l d q a q d _ j c n v m a x o n k x f _ k _ b s t q n m _ r d r % O F r : D B 2 - 1 - 0 - 0 4 ( _ m c o d b s _ s d k x _ r d % O K : D B 3 - 1 - 1 - 1 ( * b _ t r h m f b d k k v _ k k k n n r d m h m f * _ m c % 1 ( h n m h b b q n r r , k h m j h m f v h s g n s g d q c d l d s g x , k d r s d q h  d c G F b g _ h m r s g q n t f g b _ k b h t l h n m a q h c f d r * b q d _ s h m f s g d r n , b _ k k d c ’ d f f a n w – r s q t b s t q d r k d _ c h m f s n b d k k v _ k k r s h e e d m h m f _ m c q d c t b d c l _ s q h w o n q n r h s x % A q _ b b h m h d s _ k - * 0 8 8 8 : V h k k _ s r d s _ k - * 1 / / 0 : K d u d r p t d , S q d l a k _ x d s _ k - * 1 / 0 4 ( - G F l n c h  b _ s h n m _ m c c d f q _ c _ s h n m _ q d h l o n q s _ m s e _ b s n q r c t q h m f s g d _ s s _ b j n e o _ s g n f d m r n q o g x s n o g _ f n t r h m r d b s r o n r r d r r h m f b d k k v _ k k , c d f q _ c h m f d m y x l d r r t b g _ r O L D r * O F r * _ m c O K r _ r u h q t k d m b d e _ b s n q r % B _ m s t d s _ k - * 1 / / 7 : L _ k h m n u r j x d s _ k - * 1 / 0 3 ( - S g d d u h c d m b d k h m j h m f G F s n s g d c d e d m r h u d q d , r o n m r d r n e o k _ m s r h m b k t c d r s g d a q n _ c r o d b s q t l n e o _ s g n f d m q d r h r s _ m b d n q r t r b d o s h a h k h s x o g d m n s x o d r s g _ s _ q d b q d _ s d c a x _ k s d q h m f G F , l n c h e x h m f d m y x l d r h m c h e e d q d m s o k _ m s r o d b h d r % B _ m s t d s _ k - * 1 / / 7 ( - @ k s g n t f g s g d d u h c d m b d q d k _ s h m f s n G F l d s _ a n k h r l c t q h m f _ o g h c e d d c h m f h r k h l h s d c * h s h r s g n t f g s s g _ s s g d o q d r d m b d n e G F , l n c h e x h m f d m y x l d r r t b g _ r O L D r _ m c O F r * h m s g d r _ k h u _ n e _ o g h c r * b n t k c e _ b h k h s _ s d r s x k d s o d m d s q _ s h n m s g q n t f g s g d h m s d q b d k k t k _ q l _ s q h w % L b @ k k _ m _ m c @ c _ l r * 0 8 5 0 : C q d x d q _ m c B _ l o a d k k * 0 8 7 6 : L _ d s _ k - * 0 8 8 / ( - @ c c h s h n m _ k k x * a x d w o k n q h m f s g d s q _ m r b q h o s h n m _ k o q n  k d r n e @ q _ a h c n o r h r % 9 p Y a h  c n o r h r s g Y i h Y m Y ( o k _ m s r _ s s _ b j d c a x c h e e d q d m s o _ s g n f d m r _ m c o g x s n o g _ f n t r h m r d b s r * C d U n r d s _ k - % 1 / / 4 ( e n t m c s g _ s s g d N C B S H M L C S F U I C R S C P 9 R C 0 2 % 9 s N L C H 0 2 ( f d m d v _ r r o d , b h  b _ k k x t o q d f t k _ s d c c t q h m f L v y t r o d p r h b Y d e d d c h m f x d s v _ r t m b g _ m f d c c t q h m f s g d h m s d q _ b s h n m v h s g n s g d q _ s s _ b j d q r r s t c h d c - C d r o h s d s g h r u _ k t _ a k d h m e n q l _ s h n m * s g d q d r s h k k d w h r s r _ k _ b j n e c d s _ h k d c l d b g _ m h r s h b t m c d q r s _ m c h m f _ a n t s s g d q n k d n e G F c t q h m f o k _ m s , _ o g h c h m s d q _ b s h n m r - S g d e n b t r n e s g d o q d r d m s v n q j v _ r s n b g _ q _ b s d q h y d s g d c x m _ l h b r n e G F _ m c h s r l n c h e x h m f d m y x l d r c t q h m f s g d d _ q k x r s _ f d n e _ o g h c h m e d r s _ s h n m _ m c g n v s g d r d b g _ m f d r b n t k c h m fl t d m b d s g d _ o g h c r d s s k h m f _ m c e d d c h m f a d g _ u h n q - S n s g h r d m c * s g d f k n a _ k k x c h r s q h a t s d c _ o g h c L - o d p r h b Y d _ m c @ q _ a h c n o r h r v d q d t r d c _ r s g d o k _ m s , _ o g h c h m s d q _ b s h n m l n c d k - G d q d * v d r g n v s g _ s c t q h m f d _ q k x _ o g h c h m , e d r s _ s h n m * s n s _ k O L D _ b s h u h s x _ m c l d s g _ m n k d l h r r h n m h m b q d _ r d v h s g _ b n m b n l h s _ m s c d b q d _ r d h m s g d c d f q d d n e G F l d s g x k d r s d q h  b _ s h n m - D w n f d m n t r h m g h a h s h n m n e s n s _ k O L D _ b s h u h s x k d _ c r s n _ r h f m h  b _ m s c d b q d _ r d h m _ o g h c r d s s k h m f o q d e d q d m b d h m v h k c , s x o d B n k , / o k _ m s r - E t q s g d q l n q d * a x d w o k n h s h m f s g d q d r t k s r n a s _ h m d c a x C d U n r d s _ k - % 1 / / 4 ( * s g d h m g h a h s n q x _ b s h u h s x n e @ s O L D H 0 2 _ m c h s r c d e d m r h u d q n k d c t q h m f _ o g h c h m e d r s _ s h n m v d q d h r n k _ s d c _ m c b g _ q _ b s d q h y d c - C t d s n s g d l _ q j d c o q d e d q d m b d n e L - o d p r h b Y d s n r d s s k d n m o l d h 0 2 o k _ m s r * b n m b n l h s _ m s v h s g k n m f d q o g k n d l r _ o h m f d r s h n m r n m s g d r d l t s _ m s r b n l o _ q d c v h s g s g d v h k c , s x o d f d m n s x o d r * h s g _ r a d d m c d l n m , r s q _ s d c s g _ s @ s O L D H 0 2 h r _ q d r h r s _ m b d e _ b s n q c t q h m f _ o g h c b n k n , m h y _ s h n m h m @ q _ a h c n o r h r - Q D R T K S R C d s d q l h m ] s h n m n e D ] q k x H m e d r s ] s h n m R s ] f d c t q h m f I - o d p r h b P d  : q ] a h c n o r h r H m s d q ] b s h n m O q h n q s n s g d d w o d q h l d m s r _ m c _ m _ k x r h r * h m n q c d q s n c d s d q l h m d s g d s h l d r b _ k d r n e s g d d _ q k x h m e d r s _ s h n m r s _ f d r * s g d o q n o d q r _ l o k h m f s h l d e n q s g d d w o d q h l d m s r v _ r d r s _ a k h r g d c - V d c d b h c d c s n d r s _ a k h r g _ r s g d d _ q k x _ o g h c h m e d r s _ s h n m r s _ f d s g d s h l d s g _ s _ o g h c r s n n j s n o d q e n q l s g d  q r s r t r s _ h m d c o g k n d l h m f d r s h n m e q n l s g d  q r s b n m s _ b s v h s g s g d @ q _ a h c n o r h r k d _ e % _ o g h c k _ m c h m f ( - S n s g h r * v d t r d c s g d D k d b s q h b _ k O d m d s q _ s h n m F q _ o g % D O F ( s d b g m h p t d * v g h b g b q d _ s d r c h r s h m b s fl t b s t _ s h m f u n k s _ f d o _ s s d q m r q d e d q q d c s n _ r D O F v _ u d , e n q l r * v g h b g h m s t q m g _ r a d d m d w o d q h l d m s _ k k x q d k _ s d c s n c h e e d q d m s e d d c h m f o q n b d r r d r n q _ b s h u h s h d r o d q e n q l d c a x s g d h m r d b s * h m s g h r b _ r d s g d _ o g h c L - o d p r h b Y d % E h f t q d 0 @ ( - S g d D O F q d r t k s r r g n v d c s g _ s v h m f k d r r _ c t k s L - o d p r h b Y d _ o g h c r r d s s k d c n m v h k c , s x o d k d _ u d r _ m c _ b g h d u d c s g d  q r s r t r s _ h m d c o g k n d l h m f d r s h n m _ e s d q 1 6 0 - 1 5 2 3 - 7 l h m % 3 - 4 5 / - 4 g ( e q n l k _ m c h m f % E h f t q d 0 A ( - S g t r * b n m r h c d q h m f s g _ s _ o g h c r s n n j 9 4 g s n o d q e n q l s g d  q r s r t r s _ h m d c o g k n d l h m f d r s h n m % _ m c _ c c h m f 0 g s n b n u d q o n r r h a k d u _ q h _ s h n m r ( * v d d r s _ a k h r g d c _ r _ m d _ q k x h m e d r s _ s h n m r s _ f d s g d  q r s 5 g n e o k _ m s _ o g h c h m s d q _ b s h n m * _ m c a _ r d c n m s g h r s h l h m f * e t q s g d q d w , o d q h l d m s r v d q d c n m d - @ c c h s h n m _ k k x * D O F _ m _ k x r h r q d u d _ k d c s g _ s _ e s d q s g d  q r s g n r s o d m d s q _ s h n m * o d q e n q l d c 3 - 1 l h m _ e s d q k _ m c h m f * g n r s s h r r t d r v d q d o q n a d c a x _ o g h c r 9 2 4 s h l d r % 2 4 o q n a d r ( * _ m c v h s g h m s g d r d o q n a h m f d u d m s r * L - o d p r h b Y d o d q e n q l d c _ m _ u d q _ f d n e 0 3 4 l d l a q _ m d o t m b s t q d r * u h r t _ k h y d c _ r o n s d m s h _ k c q n o r % E h f t q d r 0 @ _ m c 0 A ( - L n q d n u d q * c t q h m f s g d  q r s 2 5 / l h m % 5 g ( * L - o d p r h b Y d r o d m s i t r s 7 4 - 0 l h m h m m n m o q n a h m f _ b s h u h s h d r % h - d - * v h s g s g d h q r s x k d s r n t s n e s g d g n r s o k _ m s ( % E h f t q d 0 A ( - S g d q d e n q d * s g h r b n m  q l d c s g _ s _ o g h c r o d q e n q l _ m d w g _ t r s h u d d w _ l h m _ s h n m a x b n m r s _ m s k x o q n a h m f s g d g n r s s h r r t d r c t q h m f s g d d _ q k x h m e d r s _ s h n m r s _ f d % 5 g n e o k _ m s , _ o g h c h m s d q _ b s h n m ( - D ] q k x R s ] f d n e : o g h c H m e d r s ] s h n m H m b q d ] r d r S n s ] k O L D : b s h u h s x ) L d s g ] m n k D l h r r h n m r ) ] m c : a t m c ] m b d n e C d l d s g x k d r s d q h fi d c G F B n m r h c d q h m f s g _ s O L D _ b s h u h s x _ m c s g d G F l d s g x k d r s d q h  b _ s h n m r s _ s t r g _ u d a d d m c d r b q h a d c _ r c d e d m r d , q d k _ s d c d k d l d m s r c t q h m f o _ s g n f d m _ s s _ b j % B _ m s t d s _ k - * 1 / / 7 : N r n q h n d s _ k - * 1 / / 7 : Q _ h n k _ d s _ k - * 1 / 0 0 : A d s g j d d s _ k - * 1 / 0 3 ( * h s v _ r c d b h c d c s n l d _ r t q d s g d s n s _ k O L D _ b s h u h s x _ m c h s r b n m r d p t d m s d e e d b s r n m s g d G F l d s g , x k d r s d q h  b _ s h n m c d f q d d _ m c l d s g _ m n k d l h r r h n m r % E h f t q d 1 @ ( c t q h m f s g d d _ q k x r s _ f d n e o k _ m s , _ o g h c h m s d q _ b s h n m - N t q q d r t k s r r g n v d c s g _ s s n s _ k O L D _ b s h u h s x h m b q d _ r d c 9 1 /  h m _ o g h c , h m e d r s d c k d _ u d r n e @ q _ a h c n o r h r v h s g q d r o d b s s n s g d b n m s q n k o k _ m s r % h - d - * m n m h m e d r s d c : E h f t q d 1 A ( - B n m r d p t d m s k x * s g d c d f q d d n e l d s g x k d r s d q h  b _ s h n m n e G F c d b q d _ r d c r h f m h  b _ m s k x a x 0 8  % E h f t q d 1 B ( * b n m b n l h s _ m s v h s g _ s g q d d e n k c h m b q d _ r d h m s g d l d s g _ m n k d l h r r h n m r h m s g d _ o g h c , h m e d r s d c o k _ m s r b n l o _ q d c v h s g s g d b n m s q n k b n m c h s h n m % E h f t q d 1 C ( - S n u h r t _ k h y d s g d b d k k v _ k k l n c h  b _ s h n m r s g _ s n b b t q q d c _ r _ q d r t k s n e s g d h m b q d _ r d h m O L D _ b s h u h s x * h l l t m n fl t n q d r b d m b d _ r r _ x r n m h m e d b s d c _ m c b n m s q n k @ q _ a h c n o r h r k d _ u d r v d q d o d q , e n q l d c - S g d h l l t m n fl t n q d r b d m b d _ r r _ x r s g _ s v d q d c n m d s n u h r t _ k h y d s g d G F l d s g x k d r s d q h  b _ s h n m r s _ s t r * h m r h s t * r t o o n q s s g d 0 8 0 3 S g d O k _ m s B d k k q d r t k s r n a s _ h m d c h m E h f t q d r 1 A - _ m c 1 B S g d K L 0 8 l n m n b k n m _ k _ m s h a n c x * v g h b g s _ q f d s r s g d c d l d s g x k d r s d q h  d c c n l _ h m r n e G F * r g n v d c _ c n t a k h m f n e s g d r h f m _ k h m s g d o _ q d m b g x l _ s h r r t d _ m c k n v d q d o h c d q l h r n e h m e d r s d c k d _ u d r * v h s g q d r o d b s s n s g d b n m s q n k b n m c h s h n m % E h f t q d 2 : R t o o k d l d m s _ k E h f t q d r 0 ( - _ m c 1 @ c c h s h n m _ k k x * r n l d q d o k h b _ s d r v h s g K L 0 8 _ m s h a n c x q d u d _ k d c G F c d l d s g x , k d r s d q h  b _ s h n m y n m d r k n b _ k h y d c b k n r d s n _ o g h c a n c h d r _ m c r s x k d s r % R t o o k d l d m s _ k E h f t q d 0 ( * r t f f d r s h m f s g _ s G F l n c h  b _ s h n m r b n t k c a d n b b t q q h m f _ r _ b n m r d p t d m b d n e r s x k d s o d m d s q _ s h n m s g q n t f g s g d o d b s h b l _ s q h w - N m s g d n s g d q g _ m c * _ r h f m h  b _ m s 2 /  q d c t b s h n m h m s g d r h f m _ k n e s g d K L 1 / _ m s h a n c x * v g h b g q d b n f m h y d r g h f g k x l d s g x k d r s d q h  d c G F * v _ r n a r d q u d c h m s g d _ o g h c , h m e d r s d c k d _ u d r b n l o _ q d c v h s g s g d m n m h m e d r s d c o k _ m s r % E h f t q d 2 : R t o o k d l d m s _ k E h f t q d 2 ( - @ c c h s h n m _ k k x * G F d o h s n o d r v d q d l d _ , r t q d c a x d m y x l d , k h m j d c h l l t m n r n q a d m s _ r r _ x % D K H R @ ( : g n v , d u d q * m n c h e e d q d m b d r v d q d c d s d b s d c c t q h m f _ o g h c h m e d r s _ s h n m a x s g h r l d s g n c % R t o o k d l d m s _ k E h f t q d 3 ( - S g d q d e n q d * s g d r d q d r t k s r r g n v d c s g _ s d _ q k x _ o g h c h m e d r s _ s h n m h m c t b d c _ m h m b q d _ r d h m s g d s n s _ k O L D _ b s h u h s x * v h s g s g d b n m r d p t d m s c d l d s g x k d r s d q h  b _ s h n m n e G F _ m c l d s g _ m n k q d k d _ r d - D ] q k x R s ] f d n e : o g h c H m e d r s ] s h n m H m b q d ] r d r s g d B ] k b h t l B q n r r , K h m j d c G F ] m c : k s d q r s g d S n s ] k O K : b s h u h s x N m b d G F b g _ h m r _ q d c d l d s g x k d r s d q h  d c h m b d k k v _ k k r * s g d x l _ x a d c h q d b s d c s n s v n c h e e d q d m s e _ s d r 9 % 0 ( o n k x l d q a q d _ j c n v m a x O F r _ m c . n q O K r n q % 1 ( h m s d q _ b s h n m h b _ k k x v h s g n s g d q c d l d s g x k d r s d q h  d c G F b g _ h m r s g q n t f g b _ k b h t l a q h c f d r * b q d _ s h m f s g d r n , b _ k k d c d f f a n w r s q t b s t q d r % E h f t q d 3 @ : A q _ b b h m h d s _ k - * 0 8 8 8 : V h k k _ s r d s _ k - * 1 / / 0 ( - S g d m * _ r s g d d _ q k x L - o d p r h b Y d h m e d r s _ s h n m o q n b d r r h m c t b d r G F c d d r s d q h  b _ s h n m * v g h b g n e s g d r d s v n r t a r d p t d m s r s d o r % O F . O K a q d _ j c n v m n q d f f a n w _ q q _ m f d l d m s ( b n t k c a d n b b t q q h m f h m d _ q k x _ o g h c , h m e d r s d c o k _ m s r v _ r h m u d r s h f _ s d c - S n _ b g h d u d s g h r * s n s _ k O F _ m c O K _ b s h u h s h d r v d q d l d _ r t q d c - S g d q d r t k s r q d u d _ k d c s g _ s s g d s n s _ k O F _ b s h u h s x q d l _ h m r t m b g _ m f d c % E h f t q d 3 A ( * b n m b n l h s _ m s v h s g _ r h f m h  b _ m s h m b q d _ r d h m s n s _ k O K _ b s h u h s x % E h f t q d 3 B ( * h m s g d _ o g h c , h m e d r s d c o k _ m s r v h s g q d r o d b s s n s g d b n m s q n k b n m c h s h n m - R h m b d s g d n s g d q o n r r h a k d e _ s d n e c d l d s g x k d r s d q h  d c G F h r s g d h n m b q n r r , k h m j h m f * v d u h r t _ k h y d c s g d r d d o h s n o d r a x t r h m f s g d l n m n , b k n m _ k _ m s h a n c x 1 E 3 - H m s d q d r s h m f k x * h s v _ r e n t m c s g _ s s g d d f f a n w _ q q _ m f d l d m s n e G F h r r h f m h  b _ m s k x l n q d _ a t m c _ m s h m h m e d r s d c E h f t q d 0 - C d s d q l h m _ s h n m n e R _ l o k h m f S h l d e n q D _ q k x @ o g h c H m e d r s _ s h n m R s _ f d - D O F v _ r o d q e n q l d c s n d u _ k t _ s d s g d e d d c h m f a d g _ u h n q n e L - o d p r h b Y d t r h m f @ q _ a h c n o r h r % v h k c , s x o d B n k , / ( _ r _ g n r s v h s g s g d _ h l n e c d s d q l h m h m f s g d o q n o d q r _ l o k h m f s h l d q d k _ s d c s n s g d d _ q k x h m e d r s _ s h n m r s _ f d -  : ( R b g d l _ s h b q d o q d r d m s _ s h n m n e s g d a h n k n f h b _ k _ b s h u h s h d r n e s g d _ o g h c r s x k d s h m r h c d s g d g n r s o k _ m s _ m c h s r b n q q d r o n m c h m f D O F v _ u d e n q l - S g d _ q q n v o n h m s r s n s g d o n s d m s h _ k c q n o q d k _ s d c s n s g d r s x k d s d m s q x h m s n s g d r h d u d d k d l d m s r - B * h m s d q b d k k t k _ q o q n a h m f : D 0 * o g k n d l r _ k h u _ s h n m : D 1 * o g k n d l h m f d r s h n m : O c * b d k k o t m b s t q d % o n s d m s h _ k c q n o ( : R s * r s x k d s : T d * t o o d q d o h c d q l h r : U a * u _ r b t k _ q a t m c k d -  A ( D O F u _ q h _ a k d r _ m _ k x y d c s n c d  m d s g d s h l h m f n e s g d d _ q k x _ o g h c h m e d r s _ s h n m r s _ f d - L d _ m _ m c R D v d q d b _ k b t k _ s d c e q n l m 3 1 / % 1 / h m c d o d m c d m s D O F q d b n q c h m f r ( - G n l n f _ k _ b s t q n m _ m h m O k _ m s , @ o g h c H m s d q _ b s h n m 0 8 0 4 o k _ m s r * r h m b d _ r h f m h  b _ m s s q h o k h m f h m s g d r h f m _ k n e 1 E 3 _ m s h a n c x v _ r l d _ r t q d c l _ h m k x h m s g d k n v d q d o h c d q l h r _ m c o _ q d m b g x l _ s h r r t d n e s g d _ o g h c , h m e d r s d c k d _ u d r b n l o _ q d c v h s g s g d b n m s q n k b n m c h s h n m % E h f t q d r 3 C _ m c 3 D : R t o o k d l d m s _ k E h f t q d 4 ( - S g h r r t f f d r s r s g _ s * c t q h m f d _ q k x L - o d p r h b Y d h m e d r s _ s h n m * b g _ m f d r h m G F r s q t b s t q d k d _ c s n _ m h m b q d _ r d h m s g d _ a t m c _ m b d n e a n s g c d l d s g x k d r s d q h  d c _ m c h n m b q n r r , k h m j d c G F - D w n f d m n t r L n c t k ] s h n m n e O L D : b s h u h s x ] m c L d s g ] m n k D l h r r h n m r H m  t d m b d s g d : o g h c R d s s k h m f O q d e d q d m b d S g d _ a n u d q d r t k s r q d u d _ k d c s g _ s G F l d s g x k d r s d q h  b _ s h n m r s _ s t r h r r h f m h  b _ m s k x _ k s d q d c c t q h m f s g d d _ q k x o k _ m s , _ o g h c h m s d q _ b s h n m % E h f , t q d r 1 _ m c 2 : R t o o k d l d m s _ k E h f t q d r 0 s n 2 ( - G n v d u d q * v d b _ m m n s c h r s h m f t h r g v g d s g d q s g d r d G F _ k s d q _ s h n m r b n q q d r o n m c s n _ c d , e d m r h u d l d b g _ m h r l n e s g d g n r s o k _ m s n q s n s g d b n m r d p t d m b d r n e s g d _ o g h c h m e d r s _ s h n m . e d d c h m f o q n b d r r - H m n q c d q s n f _ h m _ m h m r h f g s h m s n s g h r p t d r s h n m * h s v _ r c d b h c d c s n h m u d r s h f _ s d g n v c h e e d q d m s k d u d k r n e O L D _ b s h u h s x n e s g d g n r s o k _ m s b n t k c h m fl t d m b d s g d _ o g h c a d g _ u h n q h m s d q l r n e r d s s k h m f o q d e d q d m b d - S g d  q r s _ o o q n _ b g v _ r s n d w n f d m n t r k x l n c t k _ s d s g d s n s _ k O L D _ b s h u h s x n e v h k c , s x o d B n k , / o k _ m s r _ m c s g d m r t a i d b s s g d r d o k _ m s r s n _ e q d d b g n h b d _ r r _ x * v g h b g q d u d _ k r s g d o q d e d q d m b d n e s g d _ o g h c r s n r d s s k d n m s g d l n r s r t h s _ a k d g n r s s n d r s _ a k h r g _ m d v b n k n m x % O n b g d s _ k - * 0 8 8 7 ( - S g h r v _ r _ b g h d u d c a x h m  k s q _ s h m f n m d f q n t o n e o k _ m s r v h s g 0 l f . l K d o h f _ k k n b _ s d b g h m f _ k k _ s d % D F B F : R h f l _ , @ k c q h b g ( * v g h b g g _ r a d d m c d r b q h a d c _ r _ r o d b h  b b g d l h b _ k h m g h a h s n q n e f k n a _ k O L D _ b s h u h s x % K d v h r d s _ k - * 1 / / 7 ( - S g d m * _ r d b n m c f q n t o n e o k _ m s r v _ r h m  k s q _ s d c v h s g 0 4 t m h s r . l K n q _ m f d o d d k O L D % R h f l _ , @ k c q h b g : E h f t q d 4 @ ( - @ e s d q 0 g n e s g d h m  k s q _ s h n m o q n b d c t q d * s q d _ s d c o k _ m s r o k t r _ v _ s d q , h m  k s q _ s d c b n m s q n k f q n t o % l n b j ( v d q d r t a i d b s d c s n s g d e q d d b g n h b d _ r r _ x - S g d q d r t k s r r g n v s g _ s s q d _ s l d m s v h s g s g d b g d l h b _ k O L D h m g h a h s n q D F B F q d r t k s d c h m 9 0 /  q d c t b s h n m h m s n s _ k O L D _ b s h u h s x % E h f t q d 4 A ( b n m b n l h s _ m s v h s g _ 1 - 6 , e n k c q d c t b s h n m n e l d s g _ m n k d l h r r h n m r % E h f t q d 4 C ( b n l o _ q d c v h s g s g d h m  k s q _ s h n m b n m s q n k % l n b j ( - N m s g d n s g d q g _ m c * h m  k s q _ s h n m v h s g s g d b n l l d q b h _ k n q _ m f d o d d k O L D b n b j s _ h k h m b q d _ r d c s g d s n s _ k O L D _ b s h u h s x a x 0 4  % E h f t q d 4 A ( b n l o _ q d c v h s g s g d l n b j , h m  k s q _ s d c o k _ m s r * v g h k d l d s g _ m n k d l h r r h n m r r g n v d c m n c h e e d q d m b d r a d s v d d m a n s g b n m , c h s h n m r % E h f t q d 4 C ( - E q d d b g n h b d _ r r _ x r n m s g d r d s q d _ s d c o k _ m s r E h f t q d 1 - D _ q k x O k _ m s , @ o g h c H m s d q _ b s h n m H m b q d _ r d r F k n a _ k O L D @ b s h u h s x _ m c C d b q d _ r d r s g d C d f q d d n e G F L d s g x k d r s d q h  b _ s h n m -  : ( R b g d l _ s h b q d o q d r d m s _ s h n m n e s g d G F c d l d s g x k d r s d q h  b _ s h n m o q n b d r r - O L D h r b _ s _ k x y d c a x G F c d l d s g x k d r s d q h  b _ s h n m * _ m c s g d h q _ b s h u h s x h r q d f t k _ s d c a x s g d h q o q n s d h m _ b d n t r h m g h a h s n q % O L D H r ( -  A ( S n s _ k O L D _ b s h u h s x l d _ r t q d c _ e s d q 5 g n e L - o d p r h b Y d  @ q _ a h c n o r h r h m s d q _ b s h n m v h s g 3 , v d d j , n k c v h k c , s x o d B n k , / o k _ m s r - S n s _ k o q n s d h m d w s q _ b s r e q n l q n r d s s d k d _ u d r n e v h k c , s x o d B n k , / o k _ m s r v d q d t r d c s n l d _ r t q d f k n a _ k O L D _ b s h u h s x - U _ k t d r _ q d d w o q d r r d c _ r q d k _ s h u d O L D _ b s h u h s x _ m c m n q l _ k h y d c s n s g d _ u d q _ f d v h k c , s x o d B n k , / _ b s h u h s x % m n m h m e d r s d c ( - D q q n q a _ q r q d o q d r d m s R D e q n l m 3 2 % s g q d d h m c h u h c t _ k o k _ m s r ( -  B ( C d f q d d n e l d s g x k d r s d q h  b _ s h n m % C L ( _ e s d q 5 g n e L - o d p r h b Y d  @ q _ a h c n o r h r h m s d q _ b s h n m - D q q n q a _ q r q d o q d r d m s R D e q n l m 3 2 % s g q d d h m c h u h c t _ k o k _ m s r ( -  C ( L d s g _ m n k % L d N G ( d l h r r h n m r l d _ r t q d c _ e s d q 5 g n e L - o d p r h b Y d  @ q _ a h c n o r h r h m s d q _ b s h n m v h s g 3 , v d d j , n k c v h k c , s x o d B n k , / o k _ m s r a x e t k k d u _ o n q _ s h n m g d _ c r o _ b d f _ r b g q n l _ s n f q _ o g x - U _ k t d r b n q q d r o n m c s n o o l n e l d s g _ m n k h m 0 V K n e b n k k d b s d c s q _ m r o h q _ s h n m u _ o n q - D q q n q a _ q r q d o q d r d m s R D e q n l m 3 3 % e n t q h m c h u h c t _ k o k _ m s r ( -  A ( s n  C ( @ r s d q h r j r q d o q d r d m s r h f m h  b _ m s c h e e d q d m b d r c d s d q l h m d c a x R s t c d m s  r s s d r s % ) ) * O ; / - / / 4 ( - 0 8 0 5 S g d O k _ m s B d k k r g n v d c m n r h f m h  b _ m s c h e e d q d m b d r h m _ o g h c o q d e d q d m b d v g d m b n l o _ q d c v h s g s g d h m b q d _ r d c O L D _ b s h u h s x f q n t o n e o k _ m s r % O L D h m  k s q _ s d c ( v h s g s g d b n m s q n k b n m c h s h n m % l n b j : E h f t q d 4 B ( - H m s d q , d r s h m f k x * _ r h f m h  b _ m s q d c t b s h n m h m _ o g h c r d s s k h m f v _ r n a r d q u d c e n q s g d q d c t b d c O L D _ b s h u h s x o k _ m s r % D F B F h m  k s q _ s d c ( * r h m b d n m k x 1 /  n e s g d s n s _ k _ o g h c o n o t k _ s h n m o q d e d q q d c s g n r d o k _ m s r _ r g n r s * b n l o _ q d c v h s g 2 7  _ m c 3 1  n e s g d _ o g h c o n o t k _ s h n m s g _ s o q d , e d q q d c s n r d s s k d n m l n b j , s q d _ s d c _ m c n q _ m f d o d d k O L D , s q d _ s d c o k _ m s r * q d r o d b s h u d k x % E h f t q d 4 B ( - L n q d n u d q * h s h r j m n v m s g _ s l d s g _ m n k h r _ b q h s h b _ k u n k _ s h k d c d e d m r d r h f m _ k d l h s s d c c t q h m f o g x s n o g _ f n t r h m r d b s e d d c h m f % A _ k c v h m d s _ k - * 1 / / 5 : u n m C _ g k d s _ k - * 1 / / 5 ( * _ m c b n m r h c d q h m f s g _ s n t q q d r t k s r r g n v h m b q d _ r d c l d s g _ m n k d l h r r h n m r h m _ o g h c , h m e d r s d c o k _ m s r % E h f t q d 1 C ( * h s v _ r c d b h c d c s n h m u d r s h f _ s d g n v l d s g _ m n k d l h r r h n m r b n t k c h m fl t d m b d s g d g n r s r d s s k h m f o q d e d q , d m b d n e L - o d p r h b Y d - S n _ b b n l o k h r g s g h r _ m c a _ r d c n m l d s g _ m n k d l h r r h n m r e q n l h m e d r s d c o k _ m s r * v g h b g _ u d q _ f d c / - / 8  % u . u ( % 8 / / o o l : E h f t q d 1 C ( * _ l d s g _ m n k r n k t s h n m n e / - 0  % u . u ( v _ r o q d o _ q d c s n h m  k s q _ s d v h k c , s x o d B n k , / k d _ u d r * _ m c s g d m s g d r d o k _ m s r v d q d r t a i d b s d c s n _ m _ o g h c e q d d b g n h b d _ r r _ x t r , h m f v _ s d q , h m  k s q _ s d c o k _ m s r _ r b n m s q n k r % l n b j ( - @ r r g n v m h m E h f t q d 4 D * s g d q d r t k s r q d u d _ k d c s g _ s _ o g h c r r h f m h  b _ m s k x o q d e d q s n r d s s k d n m l d s g _ m n k , h m  k s q _ s d c o k _ m s r * r h m b d 5 /  n e s g d s n s _ k _ o g h c o n o t k _ s h n m b g n r d s g n r d o k _ m s r _ r g n r s b n l o _ q d c v h s g s g d 3 /  n e h m r d b s r s g _ s b g n r d s g d l n b j o k _ m s r - S g d r d q d r t k s r r t f f d r s s g _ s a n s g d w n f d m n t r l n c t k _ s h n m n e O L D _ b s h u h s x _ m c l d s g _ m n k d l h r r h n m h m @ q _ a h c n o r h r k d _ u d r b n t k c h m fl t d m b d s g d L - o d p r h b Y d r d s s k h m f o q d e d q d m b d - G n v d u d q * b n m r h c d q h m f s g _ s s g d h m  k s q _ s h n m o q n b d c t q d b n t k c k d _ c s n t m j m n v m b g _ m f d r h m s g d o k _ m s o g x r h n k n f x _ m c b n m r d p t d m s k x _ k s d q s g d _ o g h c a d g _ u h n q * _ r d b n m c _ o o q n _ b g v _ r c d r h f m d c h m n q c d q s n c d s d q l h m d s g d h m fl t d m b d n e s g d O L D _ b s h u h s x n u d q s g d r d s s k h m f a d g _ u h n q n e _ o g h c r - O L D H 0 2 O n r r d r r d r h m U h s q n ] m c h m U h u n H m g h a h s n q x : b s h u h s x n e O L D r ) ] m c o l d h . 1 L t s ] m s K h m d r : q d L n q d R t r b d o s h a k d s n I - o d p r h b P d R d s s k h m f D w o q d r r h n m _ m _ k x r h r t r h m f _ l h b q n _ q q _ x o t a k h r g d c a x C d U n r d s _ k - % 1 / / 4 ( r g n v d c s g _ s _ O L D h m g h a h s n q % O L D H 0 2 ( h r r o d b h  , b _ k k x t o q d f t k _ s d c c t q h m f L - o d p r h b Y d h m e d r s _ s h n m n e @ q _ a h c n o r h r - B n m r h c d q h m f s g h r  m c h m f * s g d o n s d m s h _ k q n k d n e O L D H 0 2 c t q h m f s g d o k _ m s , _ o g h c h m s d q _ b s h n m v _ r d u _ k t _ s d c - S v n S , C M @ h m r d q s h n m _ k l t s _ m s k h m d r v d q d h c d m s h  d c h m s g d k n b t r 9 s 4 f 5 1 2 5 / . N L C H 0 2 _ m c v d q d c d r h f m _ s d c _ r o l d h 0 2  0 % a _ b j f q n t m c B n k , / ( _ m c o l d h 0 2  1 % a _ b j f q n t m c V R 3 : R t o o k d l d m s _ k E h f t q d 5 @ ( - D w o q d r r h n m _ m _ k x r h r t r h m f Q S , O B Q _ m c Q S , p O B Q v d q d c n m d n m o l d h 0 2  0 _ m c o l d h 0 2  1 l t s _ m s o k _ m s r - @ l o k h  b _ s h n m n e s g d e t k k , k d m f s g b n c h m f r d p t d m b d n e N L C H 0 2 h m a n s g o l d h 0 2  0 _ m c o l d h 0 2  1 l t s _ m s k h m d r b n m  q l d c s g _ s a n s g l t s _ m s r _ q d j m n b j c n v m k h m d r * v h s g c d b q d _ r d r n e 5 4 - 4 _ m c 4 6 - 0  h m N L C H 0 2 s q _ m r b q h o s _ b b t l t k _ s h n m h m b n l o _ q h r n m v h s g s g d h q b n q q d r o n m c h m f v h k c , s x o d f d m n s x o d r * q d r o d b s h u d k x % R t o o k d l d m , s _ k E h f t q d r 5 A _ m c 5 B ( - S g d m * h m n q c d q s n b g _ q _ b s d q h y d s g d O L D , h m g h a h s h m f b _ o _ b h s x n e O L D H 0 2 * s g d h m g h a h s n q x d e e d b s n e q d , b n l a h m _ m s O L D H 0 2 n m f k n a _ k O L D _ b s h u h s x n e v h k c , s x o d o k _ m s r a x t r h m f _ f d k c h e e t r h n m _ r r _ x v _ r c d s d q l h m d c * _ r c d r b q h a d c a x R _ d y , @ f t _ x n d s _ k - % 1 / 0 2 * 1 / 0 6 ( - S g d q d r t k s r o q d r d m s d c h m R t o o k d l d m s _ k E h f t q d 5 C r g n v s g _ s s g d h m c t b d c a _ b s d q h _ k b t k s t q d b n m s _ h m h m f s g d q d b n l a h m _ m s O L D H 0 2 % O L D H 0 2 w 5 g h r  H O S F ( g _ r 2 /  _ m c 1 2  k d r r f k n a _ k O L D _ b s h u h s x s g _ m b t k s t q d r b n m s _ h m h m f s g d d l o s x u d b s n q % D U  H O S F ( _ m c s g d m n m h m c t b d c O L D H 0 2 b n m r s q t b s % O L D H 0 2 w 5 g h r “ H O S F ( * q d r o d b s h u d k x - S g t r * s g d r d q d r t k s r b n m  q l s g _ s O L D H 0 2 h r _ m h m g h a h s n q n e o d b s h m l d s g x k d r s d q _ r d _ b s h u h s x - S n b n m  q l s g d h m u h u n h m g h a h s n q _ b s h u h s x n e O L D H 0 2 h m @ q _ a h c n o r h r * s n s _ k O L D _ b s h u h s x v _ r l d _ r t q d c h m 3 , v d d j , n k c o l d h 0 2  0 _ m c o l d h 0 2  1 o k _ m s r - S g d q d r t k s r r g n v s g _ s a n s g o l d h 0 2 l t s _ m s k h m d r o n r r d r r g h f g d q s n s _ k O L D _ b s h u h s x b n l o _ q d c v h s g s g d v h k c , s x o d f d m n s x o d r - o l d h 0 2  0 r g n v d c 0 3  l n q d O L D _ b s h u h s x b n l , o _ q d c v h s g v h k c , s x o d B n k , / * v g h k d o l d h 0 2  1 d w g h a h s d c 0 0  l n q d O L D _ b s h u h s x b n l o _ q d c v h s g v h k c , s x o d V R 3 % R t o o k d l d m s _ k E h f t q d 6 @ ( - S g d r d r h f m h  b _ m s h m b q d _ r d r h m s n s _ k O L D _ b s h u h s x n a , r d q u d c h m o l d h 0 2 l t s _ m s r v d q d b n m r h r s d m s v h s g s g d h m b q d _ r d c E h f t q d 2 - D _ q k x @ o g h c H m e d r s _ s h n m H m b q d _ r d r s g d @ a t m c _ m b d n e C d l d s g x k d r , s d q h  d c G F D o h s n o d r - Q d o q d r d m s _ s h u d s q _ m r u d q r d r d b s h n m r n e 3 , v d d j , n k c v h k c , s x o d B n k , / @ q _ a h , c n o r h r k d _ u d r v d q d h l l t m n k _ a d k d c v h s g K L 0 8 _ m c K L 1 / l n m n b k n m _ k _ m s h a n c h d r s n s _ q f d s c d l d s g x k d r s d q h  d c G F % f q d d m ( _ m c g h f g k x l d s g x k , d r s d q h  d c G F % x d k k n v ( * q d r o d b s h u d k x - B _ k b n fl t n q V g h s d v _ r _ o o k h d c s n r s _ h m _ k k b d k k v _ k k r % l _ f d m s _ ( - S g d h l _ f d r r g n v b k n r d t o r n e s g d k n v d q d o h c d q l h r _ m c o _ q d m b g x l _ r t q q n t m c h m f s g d l _ h m u _ r b t k _ q a t m c k d n e k d _ u d r n e m n m h m e d r s d c _ m c h m e d r s d c o k _ m s r - K d * k n v d q d o h c d q l h r : O _ * o _ q d m b g x l _ : U a * u _ r b t k _ q a t m c k d - A _ q r 3 4 /  l - S g d f q _ o g r _ s q h f g s r g n v s g d q d k _ s h u d fl t n q d r b d m b d r h f m _ k n e d _ b g _ m s h a n c x - U _ k t d r v d q d m n q l _ k h y d c v h s g q d r o d b s s n s g d m n m h m e d r s d c b n m c h s h n m - D q q n q a _ q r q d o q d r d m s s g d R D n a s _ h m d c e q n l e n t q a h n k n f h b _ k q d o k h b _ s d r % e n t q k d _ u d r e q n l c h e e d q d m s o k _ m s r * e q n l c h e e d q d m s b t k s t q d a _ s b g d r ( - @ r s d q h r j r q d o q d r d m s r h f m h  b _ m s c h e e d q d m b d r c d s d q l h m d c a x R s t , c d m s  r s s d r s % ) * O ; / - / 4 _ m c ) ) * O ; / - / / 4 ( - G n l n f _ k _ b s t q n m _ m h m O k _ m s , @ o g h c H m s d q _ b s h n m 0 8 0 6